Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
On this passage Mr. Wallace remarks that the last five words "clearly
show that such characters are usually not 'specific,' in the sense that
they are such as distinguish species from one another, but are found in
numerous allied species." But I cannot see that the passage shows
anything of the sort. What to my mind it does show is, (_a_) that Mr.
Darwin repudiated Mr. Wallace's doctrine touching the _necessary_
utility of _all_ specific characters: (_b_) that he takes for granted
the contrary doctrine touching the inutility of _some_ specific
characters: (_c_) that without in this place alluding to the
proportional number of useless specific characters, he refers their
origin in some cases to "the nature of the organism" (i.e. "spontaneous
variability" due to internal causes), and in other cases to "the
conditions" (i.e. variability induced by external causes): (_d_) that
when established as a specific character by heredity, such a useless
character was held by him not to tend to become obsolete by the
influence of natural selection or any other cause; but, on the contrary,
to be "transmitted in nearly the same state to numerous, otherwise
modified, descendants"--or progeny of the species in genera, families,
&c.: (_e_) and, therefore, that useless characters which are now
distinctive of genera, families, &c., were held by him frequently, if
not usually, to point to uselessness of origin, when first they arose as
merely specific characters. Even the meaning which Mr. Wallace reads
into this passage must imply every one of these points; and therefore I
do not see that he gains much by apparently seeking to add this further
meaning--viz. that in Darwin's opinion there must have been some
unassignable reason preventing the occurrence of useless specific
characters in cases where species are _not_ destined to become the
parents of genera.
Public-domain text, read in full here on John Shaqi.
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