Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and UtilityRomanes, George John
Science
Darwin, and After Darwin, Volume 2 of 3: Post-Darwinian Questions: Heredity and Utility
Romanes, George John
Evolution (Biology); Heredity
"There can be no doubt, from the facts given in the early part of
this chapter, that extremely slight changes in the conditions of
life sometimes act in a definite manner on our already variable
domesticated productions [productions, therefore, with regard to
which uniformity and 'stability' of modification are least likely
to arise]; and, as the action Of changed conditions in causing
general or indefinite variability is accumulative, so it may be
with their definite action. Hence it is possible that _great_ and
_definite_ modifications of structure may result from altered
conditions acting during a long series of generations. In some few
instances a marked effect has been produced quickly on _all_, or
_nearly all_, the individuals which have been exposed to some
considerable change of climate, food, or other circumstance[172]."
[172] _Variation_, &c., vol. ii. p. 280.
Once more, in order to show that he retained these views to the end of
his life, I may quote a passage from the second edition of the _Descent
of Man_, which is the latest expression of his opinion upon these
points:--
"Each of the endless diversities in plumage, which we see in our
domesticated birds, is, of course, the result of some definite
cause; and under natural and more uniform conditions, some one
tint, _assuming that it was in no way injurious, would almost
certainly sooner or later prevail_. The free-intercrossing of the
many individuals belonging to the same species would ultimately
tend to make any change of colour thus induced _uniform in
character_.... Can we believe that the very slight differences in
tints and markings between, for instance, the female black-grouse
and red-grouse serve as a protection? Are partridges as they are
now coloured, better protected than if they had resembled quails?
Do the slight differences between the females of the common
pheasant, the Japan and golden pheasants, serve as a protection, or
might not their plumage have been interchanged with impunity? From
what Mr. Wallace has observed of the habits of certain gallinaceous
birds in the East, he thinks that such slight differences are
beneficial. For myself, I will only say, I am not convinced[173]."
[173] _Descent of Man_, pp. 473-4.
Yet "convinced" he certainly must have been on merely _a priori_
grounds, had he countenanced Mr. Wallace's reasoning from the general
theory of natural selection; and the fact that he here fails to be
convinced even by "what Mr. Wallace has observed of the habits of
certain gallinaceous birds," appears to indicate that he had considered
the question of utility with special reference to Mr. Wallace's opinion.
That opinion was then, as now, the avowed result of a theoretical
prepossession; and this prepossession, as the above quotations
sufficiently show, was expressly repudiated by Darwin.
Public-domain text, read in full here on John Shaqi.
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