Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection — John Shaqi
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
That natural selection differs from most of the other forms of isolation
in not being capable of causing _divergent_ or _polytypic_ evolution
must at once become evident, if we remember that the only way in which
isolation of any form can cause such evolution is by partitioning a
given group of intergenerants into two or more groups, each of which is
able to survive as thus separated from the other, and so to carry on the
evolution in divergent lines. But the distinguishing peculiarity of
natural selection, considered as a form of isolation, is that it effects
the isolation _by killing off all the individuals which it fails to
isolate_: consequently, this form of isolation differs from other forms
in prohibiting the possibility of any ramification of a single group of
intergenerants into two or more groups, for the purpose of carrying on
the evolution in divergent lines. Therefore, under this form of
isolation alone, evolution must proceed, palm-like, in a single line of
growth. So to speak, the successive generations continuously ascend to
higher things on the steps supplied by their own "dead selves"; but in
doing so they must climb a single ladder, no rung of which can be
allowed to bifurcate in the presence of the uniformity secured _for that
generation_ by the free intercrossing of the most fit. Even though
beneficial variations may arise in two or more directions
simultaneously, and all be simultaneously selected by survival of the
fittest, the effect of free intercrossing (in the absence of any other
form of isolation) will be to fuse all these beneficial variations into
one common type, and so to end in _monotypic_ evolution as before. In
order to secure _polytypic_ evolution, intercrossing between the
different beneficial variants which may arise must be prevented; and
there is nothing to prevent such intercrossing in the process of natural
selection _per se_. In order that the original group of intergenerants
should be divided and sub-divided into two or more groups of
intergenerants, some additional form of isolation must necessarily
supervene--when, of course, polytypic evolution will result. And, as Mr.
Gulick has shown, the conclusion thus established by deductive reasoning
is verified inductively by the facts of geographical distribution.
Public-domain text, read in full here on John Shaqi.
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