Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection — John Shaqi
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
of free intercrossing natural selection would be powerless to effect
divergent evolution. Where he went wrong was in not perceiving that
geographical isolation is not the only form of isolation. Had it
occurred to him that there may be other forms quite as effectual for the
prevention of free intercrossing, his essay could hardly have failed to
mark an epoch in the history of Darwinism. But, on account of this
oversight, he really weakened his main contention, namely, that in the
presence of free intercrossing natural selection must be powerless to
effect divergent evolution. This main contention I am now about to
re-argue. At present, therefore, we have only to observe that Wagner did
it much more harm than good by neglecting to perceive that free
intercrossing may be prevented in many other ways besides by migration,
and by the intervention of geographical barriers.
[2] _Darwinism_, p. 150.
[3] _The Darwinian Theory, and the Law of Migration_ (Eng. Trans.,
Stanford, London, 1873).
In order that we may set out with clearer views upon this matter, I will
make one or two preliminary remarks on the more general facts of
isolation as these are found to occur in nature.
In the first place, it is obvious that isolation admits of degrees: it
may be either total or partial; and, if partial, may occur in numberless
grades of efficiency. This is so manifest that I need not wait to give
illustrations. But now, in the second place, there is another general
fact appertaining to isolation which is not so manifest, and a clear
appreciation of which is so essential to any adequate consideration of
the subject, that I believe the reason why evolutionists have hitherto
failed to perceive the full importance of isolation, is because they
have failed to perceive the distinction which has now to be pointed out.
The distinction is, that isolation may be either discriminate or
indiscriminate. If it be discriminate, the isolation has reference to
the resemblance of the separated individuals to one another; if it be
indiscriminate, it has no such reference. For example, if a shepherd
divides a flock of sheep without regard to their characters, he is
isolating one section from the other indiscriminately; but if he places
all the white sheep in one field, and all the black sheep in another
field, he is isolating one section from the other discriminately. Or, if
geological subsidence divides a species into two parts, the isolation
will be indiscriminate; but if the separation be due to one of the
sections developing, for example, a change of instinct determining
migration to another area, or occupation of a different habitat on the
same area, then the isolation will be discriminate, so far as the
resemblance of instinct is concerned.
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