Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
The point, then, which in the first instance must be firmly fastened in
our minds is this:--so long as there is free intercrossing, heredity
cancels variability, and makes in favour of fixity of type. Only when
assisted by some form of discriminate isolation, which determines the
exclusive breeding of like with like, can heredity make in favour of
change of type, or lead to what we understand by organic evolution.
Now the forms of discriminate isolation, or homogamy, are very numerous.
When, for example, any section of a species adopts somewhat different
habits of life, or occupies a somewhat different station in the economy
of nature, homogamy arises within that section. There are forms of
homogamy on which Darwin has laid great stress, as we shall presently
find. Again, when for these or any other reasons a section of a species
becomes in any small degree modified as to form or colour, if the
species happens to be one where any psychological preference in pairing
can be exercised--as is very generally the case among the higher
animals--exclusive breeding is apt to ensue as a result of such
preference; for there is abundant evidence to show that, both in birds
and mammals, sexual selection is usually opposed to the intercrossing of
dissimilar varieties. Once more, in the case of plants, intercrossing of
dissimilar varieties may be prevented by any slight difference in their
seasons of flowering, of topographical stations, or even, in the case of
flowers which depend on insects for their fertilization, by differences
in the instincts and preferences of their visitors.
But, without at present going into detail with regard to these different
forms of discriminate isolation, there are still two others, both of
which are of much greater importance than any that I have hitherto
named. Indeed, these two forms are of such immeasurable importance, that
were it not for their virtually ubiquitous operation, the process of
organic evolution could never have begun, nor, having begun, continued.
Public-domain text, read in full here on John Shaqi.
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