Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
To sum up. Although in his later years, as shown by his correspondence,
Darwin came to recognize more fully the swamping effects of free
intercrossing, and the consequent importance of "separation" for the
prevention of these effects, and although in this connexion he likewise
came more clearly to distinguish between the "two cases" of monotypic
and polytypic evolution, it is evident that he never worked out any of
these matters--"thinking it prudent," as he wrote with reference to them
in 1878, "now I am growing old, to work at easier subjects[36]."
Therefore he never clearly saw, on the one hand, that free
intercrossing, far from constituting a "difficulty" to _monotypic_
evolution by natural selection, is the very means whereby natural
selection is in this case enabled to operate; or, on the other hand,
that, in the case of _polytypic_ evolution, the "difficulty" in question
is so absolute as to render such evolution, by natural selection alone,
absolutely impossible. Hence, although in one sentence of the _Origin of
Species_ he mentions three forms of isolation (besides the geographical
form) as serving in some cases to assist natural selection in causing
"divergence of character" (i. e. polytypic evolution[37]), on account of
not perceiving how great and how sharp is the distinction between the
two kinds or "cases" of evolution, he never realized that, where "two or
more new species" are in course of differentiation, _some_ form of
isolation other than natural selection must _necessarily_ be present,
whether or not natural selection be likewise so. The nearest approach
which he ever made to perceiving this necessity was in one of his
letters to Wagner above quoted, where, after again appealing to the
erroneous analogy between monotypic evolution and "unconscious
selection," he says:--"But I admit that by this process (i. e.
unconscious selection) two or more new species could hardly be formed
within the same limited area: some degree of separation, if not
indispensable, would be highly advantageous; and here your facts and
views will be of great value." But even in this passage the context
shows that by "separation" he is thinking exclusively of _geographical_
separation, which he rightly enough concludes (as against Wagner) need
certainly not be "indispensable." Had he gone a step further, he must
have seen that separation, _in some form or another, is_ "indispensable"
to polytypic evolution. Instead of taking this further step, however,
two years later he wrote to Semper as follows:--
[36] _Life and Letters_, vol. iii. p. 161.
[37] Page 81. The three forms of isolation mentioned are, "from
haunting different stations, from breeding at slightly different
seasons, or from the individuals of each variety preferring to pair
together."
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account