Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Nevertheless, while we must lament these shortcomings on the part of
Wagner, we ought to remember that he rendered important services in the
way of calling attention to the swamping effects of free intercrossing,
and, still more, in that of showing the high importance of geographical
isolation as a factor of organic evolution. Therefore, although in an
elaborate criticism of his views Weismann was easily able to dispose of
his generalizations in the imperfect form that they presented, I do not
think it was just in Weismann to remark, "if Wagner had confined himself
to the statement that geographical isolation materially assists the
process of natural selection, and thus also promotes the origination of
new species, he would have met with little or no opposition; but then,
of course, in saying this much, he would not have been saying anything
new." No doubt, as I have just shown, he _ought_ thus (as well as in
other and still more important respects not perceived by Prof. Weismann)
to have limited his statement; but, had he done so, it does not follow
that he would not have been saying anything new. For, in point of fact,
in as far as he said what was true, he did say a great deal that was
also new. Thus, most of what he said of the _principle of separation_
(apogamy) was as new as it was true, although, as we have seen, he said
it to very little purpose on account of his identifying this principle
as a whole with that of but one of its forms. Again, notwithstanding
this great error, or oversight, he certainly showed of the particular
form in question--viz. geographical isolation--that it was of
considerably _more_ importance than had previously been acknowledged.
And this was so far a valuable contribution to the general theory of
descent.
* * * * *
Prof. Weismann's essay, to which allusion has just been made[44], was,
however, in all respects a great advance upon those of Wagner. It was
not only more comprehensive in its view of the whole subject of
geographical isolation, but likewise much more adequate in its general
treatment thereof. Its principal defects, in my judgement, were, first,
the inordinately speculative character of some of its parts, and,
second, the restriction of its analysis to but one form of isolation--a
defect which it shares with the essays of Wagner, and in quite as high a
degree. Furthermore, although this essay had the great merit of
enunciating the principle of Amixia, it did so in a very inefficient
manner. For not only was this principle adduced with exclusive reference
to _geographical_ isolation, but even in regard to this one kind of
isolation it was presented in a highly inconsistent manner, as I will
now endeavour to show.
[44] _Ueber den Einfluss der Isolirung auf die Artbildung_ (1872).
Public-domain text, read in full here on John Shaqi.
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