Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
Over and over again Moritz Wagner insists, as constituting the
fundamental doctrine of his attempted reform of Darwinism, that
evolution by natural selection is impossible, unless natural selection
be assisted by geographical isolation, in order to prevent the swamping
effects of intercrossing[43]. Now, if instead of "evolution" he had said
"divergence of type," and if instead of "geographical isolation" he had
said "prevention of intercrossing," he would have enunciated the general
doctrine which it has been the joint endeavour of Mr. Gulick and myself
to set forth. But by not perceiving that "evolution" is of two radically
different kinds--polytypic and monotypic--he entirely failed to perceive
that, while for one of its kinds the _prevention_ of intercrossing is an
absolute necessity, for the other of its kinds the _permission_ of
intercrossing is a necessity no less absolute. And, again, in missing
the fact that geographical isolation is but one of the many ways
whereby intercrossing may be prevented, he failed to perceive that, even
as regards the case of polytypic evolution, he greatly erred in
representing this one form of isolation as being universally a necessary
condition to the process. The necessary condition to this process is,
indeed, the prevention of intercrossing _by some means or another_; but
his unfortunate insistence on geographical separation as the only
possible means to this end--especially when coupled with his no less
unfortunate disregard of monotypic evolution--caused him to hinder
rather than to advance a generalization which he had only grasped in
part. And this generalization is, as now so repeatedly stated, that
while the form of isolation which we know as natural selection depends
for its action upon the intercrossing of all the individuals which it
isolates (i. e. selects), when acting alone it can produce only
monotypic evolution; but that when it is supplemented by any of the
other numerous forms of isolation, it is furnished with the necessary
condition to producing polytypic evolution--and this in as many lines of
divergent change as there may be cases of this efficient separation.
[43] For instance, speaking of common, or continuous areas, he
says:--"In this case a constant variety, or new species, cannot be
produced, because the free crossing of a new variety with the old
unaltered stock will always cause it to revert to the original type;
in other words, will destroy the new form. The formation of a real
variety, which Darwin, as we know, regards as the commencement of a
new species, will only succeed when a few individuals, having
crossed the barrier of their habitat, are able to separate
themselves for a long time from the old stock." And the last
sentence, given as a summary of his whole doctrine, is--"The
geographical isolation of the form, a necessary consequence of
migration, is the cause of its typical character."
Public-domain text, read in full here on John Shaqi.
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