Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological SelectionRomanes, George John
Science
Darwin, and After Darwin, Volume 3 of 3: Post-Darwinian Questions: Isolation and Physiological Selection
Romanes, George John
Evolution (Biology); Heredity
The other most important form of discriminate isolation to which I have
alluded is Natural Selection. To some evolutionists it has seemed
paradoxical thus to regard natural selection as a form of isolation; but
a little thought will suffice to show that such is really the most
accurate way of regarding it. For, as Mr. Gulick says, "Natural
selection is the exclusive breeding of those better adapted to the
environment: ... it is a process in which the fittest are prevented from
crossing with the less fitted, by the exclusion of the less fitted."
Therefore it is, strictly and accurately, a mode of isolation, where the
isolation has reference to adaptation, and is secured in the most
effectual of possible ways--i.e. by the destruction of all individuals
whose intercrossing would interfere with the isolation. Indeed, the very
term "natural _selection_" shows that the principle is tacitly
understood to be one of isolation, because this name was assigned to the
principle by Darwin for the express purpose of marking the analogy that
obtains between it and the intentional isolation which is practised by
breeders, fanciers, and horticulturists. The only difference between
"natural selection" and "artificial selection" consists in this--that
under the former process the excluded individuals must necessarily
perish, while under the latter they need not do so. But clearly this
difference is accidental: it is in no way essential to the process
considered as a process of discriminate isolation. For, as far as
homogamous breeding is concerned, it can matter nothing whether the
exclusion of the dissimilar individuals is effected by separation or by
death.
Natural selection, then, is thus unquestionably a form of isolation of
the discriminate kind; and therefore, notwithstanding its unique
importance in certain respects, considered as a principle of organic
evolution it is less fundamental--and also less extensive--than the
principle of isolation in general. In other words, it is but a part of a
much larger whole. It is but a particular form of a general principle,
which, as just shown, presents many other forms, not only of the
discriminate, but likewise of the indiscriminate kind. Or, reverting to
the terminology of logic, it is a sub-species of the species Homogamy,
which in its turn is but a constituent part of the genus Isolation.
Public-domain text, read in full here on John Shaqi.
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