Thus we have in these apparently random forms of the calcareous bodies, complex adaptations in which every little detail as to direction, curve, and pointing is exactly determined. That they have selection-value in their present perfected form is beyond all doubt, since the animals are enabled by means of them to bore rapidly into the ground and so to escape from enemies. We do not know what the initial stages were, but we cannot doubt that the little improvements, which occurred as variations of the originally simple slimy bodies of the Holothurians, were preserved because they already possessed selection-value for the Synaptidae. For such minute microscopic structures whose form is so delicately adapted to the role they have to play in the life of the animal, cannot have arisen suddenly and as a whole, and every new variation of the anchor, that is, in the direction of the development of the two arms, and every curving of the shaft which prevented the tips from projecting at the wrong time, in short, every little adaptation in the modelling of the anchor must have possessed selection-value. And that such minute changes of form fall within the sphere of fluctuating variations, that is to say, THAT THEY OCCUR is beyond all doubt. In many of the Synaptidae the anchors are replaced by calcareous rods bent in the form of an S, which are said to act in the same way. Others, such as those of the genus Ankyroderma, have anchors which project considerably beyond the skin, and, according to Oestergren, serve "to catch plant-particles and other substances" and so mask the animal. Thus we see that in the Synaptidae the thick and irregular calcareous bodies of the Holothurians have been modified and transformed in various ways in adaptation to the footlessness of these animals, and to the peculiar conditions of their life, and we must conclude that the earlier stages of these changes presented themselves to the processes of selection in the form of microscopic variations. For it is as impossible to think of any origin other than through selection in this case as in the case of the toughness, and the "drip-tips" of tropical leaves. And as these last could not have been produced directly by the beating of the heavy rain-drops upon them, so the calcareous anchors of Synapta cannot have been produced directly by the friction of the sand and mud at the bottom of the sea, and, since they are parts whose function is PASSIVE the Lamarckian factor of use and disuse does not come into question. The conclusion is unavoidable, that the microscopically small variations of the calcareous bodies in the ancestral forms have been intensified and accumulated in a particular direction, till they have led to the formation of the anchor. Whether this has taken place by the action of natural selection alone, or whether the laws of variation and the intimate processes within the germ-plasm have cooperated will become clear in the discussion of germinal selection.