An obvious result of such partial extinction is clearly enough brought to
view The European oaks (like the American species) greatly tend to vary
that is they manifest an active disposition to produce new forms Every form
tends to become hereditary and so to pass from the state of mere variation
to that of race and of these competing incipient races some only will
survive. Quercus Robur offers a familiar illustration of the manner in
which one form may in the course of time become separated into two or more
distinct ones.
To Linnaeus this common oak of Europe was all of one species. But of late
years the greater number of European botanists have regarded it as
including three species, Q. pedunculata, Q. sessiliflora, and Q. pubescens.
De Candolle looks with satisfaction to the independent conclusion which he
reached from a long and patient study of the forms (and which Webb, Gay,
Bentham, and others, had equally reached), that the view of Linnaeus was
correct, inasmuch as it goes to show that the idea and the practical
application of the term species have remained unchanged during the century
which has elapsed since the publication of the "Species Plantarum." But,
the idea remaining unchanged, the facts might appear under a different
aspect, and the conclusion be different, under a slight and very supposable
change of circumstances. Of the twenty-eight spontaneous varieties of Q.
Robur, which De Candolle recognizes, all but six, he remarks, fall
naturally under the three sub-species, pedunculata, sessiliflora, and
pubescens, and are therefore forms grouped around these as centres; and,
moreover, the few connecting forms are by no means the most common. Were
these to die out, it is clear that the three forms which have already been
so frequently taken for species would be what the group of four or five
provisionally admitted species which closely surround Q. Robur now are. The
best example of such a case, as having in all probability occurred through
geographical segregation and partial extinction, is that of the cedar, thus
separated into the Deodar, the Lebanon, and the Atlantic cedars--a case
admirably worked out by Dr. Hooker two or three years ago. [IV-2]
A special advantage of the Cupuliferae for determining the probable
antiquity of existing species in Europe, De Candolle finds in the size and
character of their fruits. However it may be with other plants (and he
comes to the conclusion generally that marine currents and all other means
of distant transport have played only a very small part in the actual
dispersion of species), the transport of acorns and chestnuts by natural
causes across an arm of the sea in a condition to germinate, and much more
the spontaneous establishment of a forest of oaks or chestnuts in this way,
De Candolle conceives to be fairly impossible in itself, and contrary to
all experience. From such considerations, i.e., from the actual dispersion
Public-domain text, read in full here on John Shaqi.
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