of the existing species (with occasional aid from post-tertiary deposits),
it is thought to be shown that the principal Cupuliferae of the Old World
attained their actual extension before the present separation of Sicily,
Sardinia and Corsica, and of Britain, from the European Continent.
This view once adopted, and this course once entered upon, has to be
pursued farther. Quercus Robur of Europe with its bevy of admitted
derivatives, and its attending species only provisionally admitted to that
rank, is very closely related to certain species of Eastern Asia, and of
Oregon and California--so closely that "a view of the specimens by no means
forbids the idea that they have all originated from Q. Robur, or have
originated, with the latter, from one or more preceding forms so like the
present ones that a naturalist could hardly know whether to call them
species or varieties." Moreover, there are fossil leaves from diluvian
deposits in Italy, figured by Gaudin, which are hardly distinguishable from
those of Q. Robur on the one hand, and from those of Q. Douglasii, etc., of
California, on the other. No such leaves are found in any tertiary deposit
in Europe; but such are found of that age, it appears, in Northwest
America, where their remote descendants still flourish. So that the probable
genealogy of Q. Robur, traceable in Europe up to the commencement of the
present epoch, looks eastward and far into the past on far-distant shores.
Quercus Ilex, the evergreen oak of Southern Europe and Northern Africa,
reveals a similar archaeology; but its presence in Algeria leads De
Candolle to regard it as a much more ancient denizen of Europe than Q.
Robur; and a Tertiary oak, Q. ilicoides, from a very old Miocene bed in
Switzerland, is thought to be one of its ancestral forms. This high
antiquity once established, it follows almost of course that the very
nearly-related species in Central Asia, in Japan, in California, and even
our own live-oak with its Mexican relatives, may probably enough be
regarded as early offshoots from the same stock with Q. hex.
In brief--not to continue these abstracts and remarks, and without
reference to Darwin's particular theory (which De Candolle at the close
very fairly considers)--if existing species, or many of them, are as
ancient as they are now generally thought to be, and were subject to the
physical and geographical changes (among them the coming and the going of
the glacial epoch) which this antiquity implies; if in former times they
were as liable to variation as they now are; and if the individuals of the
same species may claim a common local origin, then we cannot wonder that
"the theory of a succession of forms by deviations of anterior forms"
should be regarded as "the most natural hypothesis," nor at the general
advance made toward its acceptance.
Public-domain text, read in full here on John Shaqi.
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