Having conceived the idea that varieties are incipient species, he is led
to study variation in the field where it shows itself most strikingly, and
affords the greatest facilities to investigation. Thoughtful naturalists
have had increasing grounds to suspect that a reexamination of the question
of species in zoology and botany, commencing with those races which man
knows most about, viz., the domesticated and cultivated races, would be
likely somewhat to modify the received idea of the entire fixity of
species. This field, rich with various but unsystematized stores of
knowledge accumulated by cultivators and breeders, has been generally
neglected by naturalists, because these races are not in a state of nature;
whereas they deserve particular attention on this very account, as
experiments, or the materials for experiments, ready to our hand. In
domestication we vary some of the natural conditions of a species, and thus
learn experimentally what changes are within the reach of varying conditions
in Nature. We separate and protect a favorite race against its foes or its
competitors, and thus learn what it might become if Nature ever afforded it
equal opportunities. Even when, to subserve human uses, we modify a
domesticated race to the detriment of its native vigor, or to the extent of
practical monstrosity, although we secure forms which would not be
originated and could not be perpetuated in free Nature, yet we attain wider
and juster views of the possible degree of variation. We perceive that some
species are more variable than others, but that no species subjected to the
experiment persistently refuses to vary; and that, when it has once begun
to vary, its varieties are not the less but the more subject to variation.
"No case is on record of a variable being ceasing to be variable under
cultivation." It is fair to conclude, from the observation of plants and
animals in a wild as well as domesticated state, that the tendency to vary
is general, and even universal. Mr. Darwin does "not believe that
variability is an inherent and necessary contingency, under all
circumstances, with all organic beings, as some authors have thought." No
one supposes variation could occur under all circumstances; but the facts
on the whole imply a universal tendency, ready to be manifested under
favorable circumstances. In reply to the assumption that man has chosen for
domestication animals and plants having an extraordinary inherent tendency
to vary, and likewise to withstand diverse climates, it is asked:
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