Geology makes huge demands upon time; and we regret to find that it has
exhausted ours--that what we meant for the briefest and most general sketch
of some geological considerations in favor of Darwin's hypothesis has so
extended as to leave no room for considering "the great facts of
comparative anatomy and zoology" with which Darwin's theory "very well
accords," nor for indicating how "it admirably serves for explaining the
unity of composition of all organisms, the existence of representative and
rudimentary organs, and the natural series which genera and species
compose." Suffice it to say that these are the real strongholds of the new
system on its theoretical side; that it goes far toward explaining both the
physiological and the structural gradations and relations between the two
kingdoms, and the arrangement of all their forms in groups subordinate to
groups, all within a few great types; that it reads the riddle of abortive
organs and of morphological conformity, of which no other theory has ever
offered a scientific explanation, and supplies a ground for harmonizing the
two fundamental ideas which naturalists and philosophers conceive to have
ruled the organic world, though they could not reconcile them; namely,
Adaptation to Purpose and Conditions of Existence, and Unity of Type. To
reconcile these two undeniable principles is the capital problem in the
philosophy of natural history; and the hypothesis which consistently does
so thereby secures a great advantage.
We all know that the arm and hand of a monkey, the foreleg and foot of a
dog and of a horse, the wing of a bat, and the fin of a porpoise, are
fundamentally identical; that the long neck of the giraffe has the same and
no more bones than the short one of the elephant; that the eggs of Surinam
frogs hatch into tadpoles with as good tails for swimming as any of their
kindred, although as tadpoles they never enter the water; that the
Guinea-pig is furnished with incisor teeth which it never uses, as it sheds
them before birth; that embryos of mammals and birds have branchial slits
and arteries running in loops, in imitation or reminiscence of the
arrangement which is permanent in fishes; and that thousands of animals and
plants have rudimentary organs which, at least in numerous cases, are
wholly useless to their possessors, etc., etc. Upon a derivative theory
this morphological conformity is explained by community of descent; and it
has not been explained in any other way.
Naturalists are constantly speaking of "related species," of the "affinity"
of a genus or other group, and of "family resemblance"--vaguely conscious
that these terms of kinship are something more than mere metaphors, but
unaware of the grounds of their aptness. Mr. Darwin assures them that they
have been talking derivative doctrine all their lives--as M. Jourdain
talked prose--without knowing it.
Public-domain text, read in full here on John Shaqi.
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