If it is difficult and in many cases practically impossible to fix the
limits of species, it is still more so to fix those of genera; and those of
tribes and families are still less susceptible of exact natural
circumscription. Intermediate forms occur, connecting one group with
another in a manner sadly perplexing to systematists, except to those who
have ceased to expect absolute limitations in Nature. All this blending
could hardly fail to suggest a former material connection among allied
forms, such as that which the hypothesis of derivation demands.
Here it would not be amiss to consider the general principle of gradation
throughout organic Nature--a principle which answers in a general way to the
Law of Continuity in the inorganic world, or rather is so analogous to it
that both may fairly be expressed by the Leibnitzian axiom, Natura non agit
saltatim. As an axiom or philosophical principle, used to test modal laws
or hypotheses, this in strictness belongs only to physics. In the
investigation of Nature at large, at least in the organic world, nobody
would undertake to apply this principle as a test of the validity of any
theory or supposed law. But naturalists of enlarged views will not fail to
infer the principle from the phenomena they investigate--to perceive that
the rule holds, under due qualifications and altered forms, throughout the
realm of Nature; although we do not suppose that Nature in the organic
world makes no distinct steps, but only short and serial steps--not
infinitely fine gradations, but no long leaps, or few of them.
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