Darwiniana : Essays — Volume 02Huxley, Thomas Henry
Science
Darwiniana : Essays — Volume 02
Huxley, Thomas Henry
Darwin, Charles, 1809-1882; Evolution (Biology)
Thus while it may be doubted if, for some years, any one is likely to be
competent to pronounce judgment on all the issues raised by Mr. Darwin,
there is assuredly abundant room for him, who, assuming the humbler, though
perhaps as useful, office of an interpreter between the "Origin of Species"
and the public, contents himself with endeavouring to point out the nature
of the problems which it discusses; to distinguish between the ascertained
facts and the theoretical views which it contains; and finally, to show the
extent to which the explanation it offers satisfies the requirements of
scientific logic. At any rate, it is this office which we purpose to
undertake in the following pages.
It may be safely assumed that our readers have a general conception of the
nature of the objects to which the word "species" is applied; but it has,
perhaps, occurred to a few, even to those who are naturalists _ex
professo_, to reflect, that, as commonly employed, the term has a double
sense and denotes two very different orders of relations. When we call a
group of animals, or of plants, a species, we may imply thereby, either
that all these animals or plants have some common peculiarity of form or
structure; or, we may mean that they possess some common functional
character. That part of biological science which deals with form and
structure is called Morphology--that which concerns itself with function,
Physiology--so that we may conveniently speak of these two senses, or
aspects, of "species"--the one as morphological, the other as
physiological. Regarded from the former point of view, a species is nothing
more than a kind of animal or plant, which is distinctly definable from all
others, by certain constant, and not merely sexual, morphological
peculiarities. Thus horses form a species, because the group of animals to
which that name is applied is distinguished from all others in the world by
the following constantly associated characters. They have--1, A vertebral
column; 2, Mammae; 3, A placental embryo; 4, Four legs; 5, A single
well-developed toe in each foot provided with a hoof; 6, A bushy tail; and
7, Callosities on the inner sides of both the fore and the hind legs. The
asses, again, form a distinct species, because, with the same characters,
as far as the fifth in the above list, all asses have tufted tails, and
have callosities only on the inner side of the fore-legs. If animals were
discovered having the general characters of the horse, but sometimes with
callosities only on the fore-legs, and more or less tufted tails; or
animals having the general characters of the ass, but with more or less
bushy tails, and sometimes with callosities on both pairs of legs, besides
being intermediate in other respects--the two species would have to be
merged into one. They could no longer be regarded as morphologically
distinct species, for they would not be distinctly definable one from the
other.
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