Darwinism (1889): An exposition of the theory of natural selection, with some of its applicationsWallace, Alfred Russel
Science
Darwinism (1889): An exposition of the theory of natural selection, with some of its applications
Wallace, Alfred Russel
Evolution; Natural selection
Another curious example of the way in which species have been modified
to occupy new places in nature, is afforded by the various animals which
inhabit the water-vessels formed by the leaves of many epiphytal species
of Bromelia. Fritz Müller has described a caddis-fly larva which lives
among these leaves, and which has been modified in the pupa state in
accordance with its surroundings. The pupae of caddis-flies inhabiting
streams have fringes of hair on the tarsi to enable them to reach the
surface on leaving their cases. But in the species inhabiting bromelia
leaves there is no need for swimming, and accordingly we find the tarsi
entirely bare. In the same plants are found curious little Entomostraca,
very abundant there but found nowhere else. These form a new genus, but
are most nearly allied to Cythere, a marine type. It is believed that
the transmission of this species from one tree to another must be
effected by the young crustacea, which are very minute, clinging to
beetles, many of which, both terrestrial and aquatic, also inhabit the
bromelia leaves; and as some water-beetles are known to frequent the
sea, it is perhaps by these means that the first emigrants established
themselves in this strange new abode. Bromeliae are often very abundant
on trees growing on the water's edge, and this would facilitate the
transition from a marine to an arboreal habitat. Fritz Müller has also
found, among the bromelia leaves, a small frog bearing its eggs on its
back, and having some other peculiarities of structure. Several
beautiful little aquatic plants of the genus Utricularia or bladder-wort
also inhabit bromelia leaves; and these send runners out to neighbouring
plants and thus spread themselves with great rapidity.
_The Importance of Isolation._
Isolation is no doubt an important aid to natural selection, as shown by
the fact that islands so often present a number of peculiar species; and
the same thing is seen on the two sides of a great mountain range or on
opposite coasts of a continent. The importance of isolation is twofold.
In the first place, it leads to a body of individuals of each species
being limited in their range and thus subjected to uniform conditions
for long spaces of time. Both the direct action of the environment and
the natural selection of such varieties only as are suited to the
conditions, will, therefore, be able to produce their full effect. In
the second place, the process of change will not be interfered with by
intercrossing with other individuals which are becoming adapted to
somewhat different conditions in an adjacent area. But this question of
the swamping effects of intercrossing will be considered in another
chapter.
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