These essays are profoundly interesting, and will supply food for
thought for many years, but it must be admitted that Weismann has
gradually been led away to speculations of the most elaborate kind,
built upon the most flimsy substratum of fact. There is evidence
of this tendency even in his early writings, but in his later
essays, especially his recent work on “The Germ Plasm: a Theory of
Heredity,”[10] the speculative part quite overpowers the rest.
As those who are interested in heredity will probably read his works
with the greatest attention, I have ventured in an appendix to
make clear what in his works may, in my opinion, safely be looked
upon as speculative rather than legitimate, or even provisional,
generalisation, and what, therefore, may be altogether omitted from
the study of a practical problem such as that with which we are
concerned.
_Are Acquired Characters transmitted?_
The practical issues which both Galton and Weismann have raised
cannot, however, be underestimated, and, in respect to the
non-inheritance of acquired characters, the mass of modern thinkers
may already be said to have given their allegiance to the views of
those two thinkers.
But we are living in times when mere authority is at a discount,
and we may well demand the facts for ourselves. The point which
we are inclined to question is one as to which a doubt was often
present in Darwin’s mind. Granted that selection is a factor, we are
inclined to believe that the transmission of acquired characters
must also take place, at any rate, to some extent. In attempting
to decide this question upon the facts themselves we may take two
lines of research. In the first place, we may examine every case
of racial change, the production of new or different parts, the
development of a new instinct, or the degeneration or loss of parts
or instincts present at some past epoch. If in every case we are
not compelled to exclude natural selection, and if in every case
that we can directly and experimentally follow, selection is the
outstanding factor, then there is strong presumptive evidence that
racial change is caused by selection and not by the inheritance
of acquired characters. In the second place, we may artificially
induce the acquisition of some character, and notice whether this
is transmitted; if it is not, then the general operation of this
kind of transmission is rendered very doubtful.
_Many Cases of Supposed Transmission to be explained by Selection._
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