Degeneration: A Chapter in DarwinismLankester, E. Ray (Edwin Ray), Sir
Science
Degeneration: A Chapter in Darwinism
Lankester, E. Ray (Edwin Ray), Sir
Degeneration; Evolution (Biology)
[Illustration: FIG. 18.—Ascidian Tadpole with a part only of the tail
_C_. _N_, nervous system with the enlarged brain in front and the narrow
spinal cord behind _n_; _N´_, is placed in the cavity of the brain:
_O_, the single cerebral eye lying in the brain; _a_, similarly placed
auditory organ; _K_, pharynx; _d_, intestine; _o_, rudiment of the mouth;
_ch_, notochord or primitive backbone. (From Gegenbaur’s “Elements of
Comparative Anatomy.”)]
It is clear then that the Ascidians must be admitted to be Vertebrates,
and must be classified in that great sub-kingdom or branch of the animal
pedigree. The Ascidian tadpole is very unlike its parent the Ascidian,
and has to go through a process of _degeneration_ in order to arrive
at the adult structure. The diagrams which are reproduced in Figs. 19
and 20, show how this degeneration proceeds. It will be observed, that
in somewhat the same manner as the young barnacle, the young Ascidian
fixes itself to a stone by its head: then the tail with its notochord
and nerve-chord atrophies. The body grows and gradually changes its
shape, whilst the cloacal chamber forms. The brain remains quite small
and undeveloped, and the remarkable myelonic eye (the eye in the brain)
disappears. The number of gill-slits increases as the animal grows in
size and its outer skin becomes tough and leather-like.
[Illustration: FIG. 19.—Degeneration of Ascidian Tadpole to form the
adult. The black pieces represent the rock or stone to which the Tadpole
has fixed its head.]
[Illustration: FIG. 20.—Very young Ascidian with only two gill-slits.
Compare with Fig 15; which is, however, seen from the other side, so that
left there corresponds to right here.]
[Illustration: FIG. 21.—Section through the eye (“surface-eye”) of a
Water-beetle’s larva. All the cells are seen to be in a row continuous
with _h_, the cells of the outermost skin or ectoderm. _p_, pigmented
cells; _r_, retinal cells connected at _o_ with the optic nerve; _g_,
transparent cells (forming a kind of “vitreous body”); _l_, cuticular
lens. (From Gegenbaur’s “Elements of Comparative Anatomy,” after
Grenacher.)]
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