Descriptions of New Hylid Frogs From Mexico and Central AmericaDuellman, William Edward
Science
Descriptions of New Hylid Frogs From Mexico and Central America
Duellman, William Edward
Frogs -- Central America; Frogs -- Mexico; Hylidae
_Variation._--The snout-vent length in seven adult males is 47.2-50.0
mm., and in five females, 45.1-52.5 mm. In neither sex do the average
proportions differ noticeably from those of the holotype, except that
the tympanum is relatively larger in females. The ratio of the diameter
of the tympanum to that of the eye is 0.363-0.468 (mean 0.438) in males
and 0.500-0.545 (mean 0.516) in females. The average number of
prevomerine teeth in males is 7.9, in females 8.4.
In life dorsal coloration varied from pale green to olive-green with
darker green or black flecks or reticulations, or pinkish tan to brown
with dark brown flecks or reticulations. Some preserved specimens have
relatively few dark flecks, whereas in most specimens the dorsum is
heavily marked. All specimens have some white markings above the anus
and on the posterodorsal surfaces of the thighs, but in some individuals
the white flecks are expanded and interconnected forming an irregular
white line.
Juveniles have a notably different coloration in life. The dorsum is
uniform pale green; the anterior and posterior surfaces of the thighs,
fingers, first three toes, and webbing are deep yellow. The anal stripe
is creamy white, and the flanks are pale gray with black flecks. The
upper lip, supratympanic fold, and canthal stripe are a bronze color.
The belly is pale yellow with a silvery cast on the throat. Juveniles
having snout-vent lengths from 24.5 to 36.6 mm. are so colored in life,
and uniform dark bluish gray dorsally in preservative.
_Comparisons._--The absence of a quadratojugal and the presence of a
greatly enlarged, non-projecting prepollex place _Hyla siopela_ in the
_Hyla bistincta_ group (see Duellman, 1964, and Adler, 1965). The
presence of a rostral keel separates _Hyla siopela_ from other members
of the _Hyla bistincta_ group, which is composed of two species having
long anal sheaths (_bistincta_ and _pentheter_), two small species
having axillary membranes and lacking nuptial excrescences in breeding
males (_charadricola_ and _chryses_), and three species (_crassa_,
_pachyderma_, and _robertsorum_) having short heads, round snouts, short
anal sheaths, and nuptial excrescences in breeding males. _Hyla siopela_
differs from the last three species in the shape of the snout and from
each in certain structural features; _H. crassa_ has fully webbed feet;
_H. pachyderma_ has large nuptial spines, and _H. robertsorum_ has more
webbing and a shorter tarsal fold. Furthermore, the venter in _H.
robertsorum_ is brown with creamy white flecks.
In structure and coloration _H. arborescandens_ resembles _siopela_, but
the former is smaller, and males of _arborescandens_ have vocal slits.
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