Disease in plantsWard, H. Marshall (Harry Marshall)
Science
Disease in plants
Ward, H. Marshall (Harry Marshall)
Plant diseases
The fact that the root-hair contains living protoplasm, enables us to
understand to a certain extent the results of the following
experiments.
If we have a leafy and healthy plant, with roots, bearing numerous
root-hairs, properly established in suitably moist soil in the pot, the
roots cease to absorb water if the temperature of the soil falls below a
certain minimum, though they recommence to do so if the temperature is
raised again: this has nothing to do with the temperature of the upper
parts of the plant, or of the air, and the latter may be so high that
the plant rapidly droops from loss of water at the leaves, which is not
being compensated owing to the inactivity of the roots.
Similarly we may have the air so cold, at a time when the soil is warm
enough to keep the root-hairs actively at work, that the plant becomes
surcharged with water, which escapes from the leaves like drops of dew.
The temperatures necessary to cause these disturbances in the action of
the living root-hairs vary for different plants, and even for different
varieties of the same species.
Similar arrestation of the functions of the roots may be brought about
by removing the oxygen from the soil around the root-hairs, and
replacing it by carbon-dioxide, or the vapour of chloroform. If not kept
too long in such a condition, the plant recovers rapidly on admitting
atmospheric oxygen, which is always present in a normal well-drained
soil both as gas in the capillary interspaces, and dissolved in the
water on the surfaces of the particles. If the access of oxygen is
delayed, however, as often happens in rainy seasons and in wet soils,
the root-hairs are killed, and rot sets in. A good instance of this has
lately been given by Heinricher in the case of potatoes.
NOTES ON CHAPTER V.
For the further pursuit of this subject the reader should
consult Sachs' _Lectures_, II. and XV.; Sorauer, _A Popular
Treatise on the Physiology of Plants_, 1895, chapters II. and
IV., and Pfeffer's _Physiology_, pp. 149-163. The principal
paper on root-hairs referred to in the text is Schwarz, "Die
Wurzelhaare der Pflanzen," in _Unters. aus dem bot. Inst. zu
Würzburg_, I. Heft 2, 1883, p. 140, where a very exhaustive
account of these organs will be found.
CHAPTER VI.
THE FUNCTIONS OF ROOT-HAIRS.
_Excretions from root-hairs--Osmotic phenomena--Turgescence--
Plasmolysis--Control of the protoplasm in absorption, etc.
Selective absorption._
We see then that the root-hairs are the active living instruments in
absorbing the water (containing small quantities of dissolved
substances) of the soil.
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