Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
Having completed the survey of the central and southern land masses
of the globe we next direct attention to the northern land masses,
viz. North and Central America on the one hand, and Europe and
northern Asia on the other. With regard to America we find besides
many species of Lumbricidae the genera _Notiodrilus_, _Microscolex_
among the Acanthodrilinae, _Megascolides_ and _Plutellus_ of the
Megascolecinae, no member at all of the Octochaetinae, a considerable
number of species of _Ocnerodrilus_ and sub-genera belonging to the
Ocnerodrilinae, a good many species of _Dichogaster_ and at least
one of _Trigaster_ among the Trigastrinae, while one sub-family,
that of the Diplocardiinae, is only found here and contains two
genera _Diplocardia_ and _Zapotecia_ with quite ten species between
them of which only one belongs to the last-named genus. There are
no Geoscolecids (except the chiefly aquatic _Sparganophilus_); this
family stops short in the West Indies where their presence has been
briefly referred to in considering the worms of South America. The
vast majority of the genera enumerated here are only found in the
warmer parts of the North American continent. We have therefore
in this division of the world some nine genera of which at least
one, viz. _Diplocardia_ (and its close ally, hardly perhaps to be
separated, _Zapotecia_), is confined to it, while _Trigaster_ only
extends as far south as to the West Indies.
Europe and northern Asia, of which Europe alone and certain
limited tracts of Asia are at all known, contain all the genera
of the family Lumbricidae which are (according to Michaelsen)
_Eiseniella_, _Eisenia_, _Helodrilus_, _Octolasium_, _Lumbricus_
with various sub-genera of _Helodrilus_, such as _Dendrobaena_ and
_Allolobophora_. Besides these forms, which amount to at least 130
species, we find the genus _Hormogaster_, with two species, the only
genus of the Geoscolecid sub-family Hormogastrinae; there is also
a species apiece of the genera _Criodrilus_ and _Sparganophilus_
(sub-family Criodrilinae). But as these are at least largely aquatic
they come under another set of rules and are not important in the
present survey of the earthworms of the world. It will be thus seen
that with these two last exceptions the area in question contains but
six genera of which _all_ are peculiar.
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