Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
of roundish sacs, with two or three minute diverticula, known as the
spermathecae. In the diverticula of these sacs are stored the sperm
derived from another individual.
[Illustration: Fig. 6. Genital region of _Maoridrilus dissimilis_
(in which species the different organs are essentially as in
_Notiodrilus tamajusi_). The oesophagus is cut away between the
Xth and XVIIIth segments to display the various organs. _n_
external orifices of nephridia which alternate in different
segments; the one lettered opens in front of dorsalmost pair of
setae, that of segment XVI opens in front of ventralmost pair,
_p_ coiled glands opening on to XVIth and XIXth segments, _sc._
sacs containing long seta, associated with these glands, _t_
spermary of segment; another pair in an exactly similar position
in segment XI. Behind spermaries are funnel-shaped openings
of sperm ducts which are seen running along the body to their
external orifice on XVIIIth segment. _v.s._ sperm sacs behind the
posterior of these and in segment XIII is seen the large ovary
corresponding in position to the spermary and opposite to each
ovary the oviduct. (x 3.)]
[Illustration: Fig. 7. A spermatheca of _Notiodrilus tamajusi_.
The external orifice is shown at _o_; above this are the
diverticula, _s_ is a portion of an intersegmental septum. (After
Eisen.)]
This completes the general sketch of the structure of _Notiodrilus
tamajusi_ which we have selected as a type. In this same genus are a
large number of species which differ from that selected in various
small structural points. Thus in _N. annectens_ (Beddard), a species
from New Zealand, the spermaries and ovaries are attached to the
posterior, instead of to the anterior, wall of their segments, and
there are neither calciferous glands nor modified setae upon segments
XVII and XIX. In all essentials however the two types agree and are
thus to be looked upon as referable to the same genus. Starting from
the structure of these types we may now sketch in quite a brief way
the main divergencies of structure shown in the group of Oligochaeta.
We shall naturally begin with the family Megascolecidae of which a
type has just been described.
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