Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
We have now to consider the eastern region of the world comprising
the two regions known generally to zoogeographers as the Oriental
and Australian. Taking a large view of the range of sub-families
and genera, and endeavouring to make the great regions of the globe
more or less equal, it seems difficult to divide further a region
which shall include all of this vast territory, and which may
therefore be termed Indo-Australian. For we find as characteristic
of the entire stretch of country the great majority of the genera
of the huge family Megascolecidae. Indeed the largest sub-family of
this family, _i.e._ the Megascolecinae, is, save for the mysterious
occurrence of the genera _Plutellus_ and _Megascolides_ in America,
absolutely limited to this area. Another sub-family, that of the
Octochaetinae, is limited to it. So far as concerns the others of
the sub-families of Megascolecidae it is only the Trigastrinae
which occur here (the genus _Eudichogaster_ and a few possibly
introduced species of _Dichogaster_) and a scattered species or two
of _Notiodrilus_ of the sub-family Acanthodrilinae. Again there are a
few and probably introduced species of the sub-family Ocnerodrilinae.
More important still this region has confined to itself the family
Moniligastridae; for a species described some years ago by myself
from the Bahamas is doubtless an introduced form. We have a complete
absence of indigenous Lumbricidae and Geoscolecidae excepting the
aquatic _Glyphidrilus_ of the sub-family Microchaetinae. It is true
that by taking isolated tracts, even large tracts, of this great
regional expanse a sub-division into well characterised regions can
be apparently recognised. But in taking such a step we shall be
confronted with the curious fact that it is rather neighbouring than
widely remote sub-divisions which present the greater differences.
If we compare for example India and New Zealand we find in common
such striking genera as _Octochaetus_, _Hoplochaetella_ and
_Diporochaeta_; whereas these genera are absent from the intervening
islands of the great Malay archipelago. On the other hand Australia
differs from the comparatively neighbouring islands of Borneo
and others by the absence in those islands of the characteristic
Australian genera such as _Megascolex_, _Notoscolex_, _Plutellus_
etc. which are in their turn found in India. It is facts like these
which render very difficult the apportioning of the tracts of country
forming the eastern hemisphere into separate regions.
There is no doubt that the Malay archipelago and the adjacent coasts
of Asia up to Japan differ from both India and Australia by the
almost entire limitation of the genus _Pheretima_ to them; but we
cannot intercalate a region in the middle of another geographical
area in this fashion!
Public-domain text, read in full here on John Shaqi.
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