Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
Having therefore arrived at a mapping out of the world into regions
in accord with its earthworm fauna, it is desirable to ascertain
what light the facts throw upon the geological and evolutionary
questions with which the study of zoogeography deals. The existence
of an antarctic region binding together such distant points as South
Georgia, the Cape of Good Hope and Kerguelen Island, seems to argue
strongly for the former extensions northwards of the antarctic
continent so far north as to embrace these several regions of that
hemisphere. In view of the facts relating to the danger of sea water
to earthworms, to their lack of facilities for migration, other than
unassisted locomotion, points which have been dealt with earlier, it
is difficult to explain their range in the antarctic hemisphere on
other grounds. The very fact that the actual earthworm fauna of New
Zealand has led us on the whole to assign it to the Indo-Australian
regions shows the inherent uselessness of the current view of
zoogeography. For were we to leave the matter here the relationship
of New Zealand to the regions of the world which lie to the south of
it would not be apparent. However, here as in so many cases there is
an antagonism between cut and dried systems and the indications of
evolution.
This assumed existence of a former antarctic continent which
connected Southern Africa and Southern America as well as various
islands has perhaps a further justification in the distribution of
the Geoscolecidae. This family is divisible into two well-marked
sub-families of which one as has already been mentioned is limited
to South America and another practically to Africa (the exceptions
being species of the largely aquatic _Glyphidrilus_), while a third
sub-family the Criodrilinae is more widely distributed--again in
accordance, one may perhaps assume, with its largely aquatic mode of
life. It is also conceivable that the genus _Dichogaster_ is another
example pointing the same way. The arguments for regarding this genus
as an indigene of the East are not strong. But there is on the
other hand no doubt that the Indian _Eudichogaster_ is very closely
allied to it. But it is by no means excluded from this argument to
suppose that these Trigastrinae owe their likeness to convergence.
At any rate there are examples of equally marked convergence which
seem to be as nearly proved as can be in another though allied
group. The New Zealand _Neodrilus_ is to all intents and purposes a
_Maoridrilus_ in which one of the two pairs of spermiducal glands
and spermathecae has disappeared. It retains the characteristic
alternation in the position of the nephridia of _Maoridrilus_, and
other structural similarities unite the two genera. In the same way
species of _Microscolex_ seem as easily derivable from _Notiodrilus_.
_Microscolex_ and _Neodrilus_ are so near that had we no such hint of
their origin it would be reasonable to place them in the same genus.
Public-domain text, read in full here on John Shaqi.
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