Earthworms and Their AlliesBeddard, Frank E. (Frank Evers)
Science
Earthworms and Their Allies
Beddard, Frank E. (Frank Evers)
Earthworms
They at least show a marked convergence.
It will be noticed therefore that the facts of their distribution
agree, as it would appear, with the structure of the terrestrial
Oligochaeta. The primitive characters of the genus _Notiodrilus_ are
to be seen in the double spermaries and glands appended to the duct,
and the corresponding spermatheca, in the absence, or very slight
development, of the papillae, so frequent in more specialised genera
such as _Pheretima_, and in the general simplicity of many organs of
the body which are more complicated elsewhere. As one would expect
with an archaic form this genus is widely ranging, being found in all
the principal land masses of the globe except in the Euro-Asiatic
continent.
Furthermore geographical facts would at least be not contradictory to
the view that this genus, and therefore the terrestrial Oligochaeta
generally, originated in the Antarctic hemisphere and that in pushing
northwards it has given off various descendants which survive in the
various regions of the world. Basing our views of the possibilities
of range among earthworms on the actual facts already dealt with, it
would seem that the peopling of America from Africa or of Africa from
America, if it has occurred, has not taken place through Europe and
the north generally. For otherwise we should expect traces of the
passage. It is true that we actually have _Hormogaster_ as a possible
sign that the Geoscolecidae have passed this way. But that is an
isolated case and may be referred to the extension northwards of this
particular genus rather than as an indication of a whole migration
through those territories. Another conclusion which a collocation of
the various facts brought together in this book appears to lead to is
that the group of the terrestrial Oligochaeta is relatively speaking
a modern one.
Convinced as we must be of the fact that range is only possible by
unaided locomotion through continuous land areas, the fact that but
few gaps occur in the range of a particular sub-family or lesser
group seems to indicate that no great time has elapsed since the
specialisation of these different forms. The dependence of earthworms
upon vegetable mould also points in the same direction and furnishes
an argument for the belief that these animals only greatly increased
on the advent of abundant dicotyledonous plants, and perhaps indeed
were actually contemporaneous with them.
LIST OF LITERATURE REFERRING TO EARTHWORMS
In the list given below I am only able to mention a few of the larger
works relating to this group. To give anything like a complete list
would demand many pages of titles. From the works selected the reader
can, if it be desired, find his way to the remaining literature of
the group.
A. GENERAL WORKS
Vejdovsky. System und Morphologie der Oligochaeten. Prag, 1884.
Beddard. A Monograph of the Oligochaeta. Oxford, 1895.
Michaelsen. Oligochaeten in 'Das Thierreich.' Berlin, 1900.
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