Ecological Studies of the Timber Wolf in Northeastern MinnesotaMech, L. David
Science
Ecological Studies of the Timber Wolf in Northeastern Minnesota
Mech, L. David
Deer -- Minnesota; Gray wolf -- Behavior -- Minnesota; Gray wolf -- Ecology -- Minnesota
It is true that wolves stand even shorter than deer and so might be
expected to flounder even more. However, this is where another factor
becomes important, the "weight-load-on-track" or total weight per area
of track. As Kelsall (1969) has pointed out, the mean
weight-load-on-track for deer is extremely difficult to measure
directly, because the actual under-surface of the deer's foot slants
vertically, and a much greater area may be used to support an animal in
snow than on a hard surface. This probably explains the discrepancy
between Kelsall's measurements and work done by Verme (1968) in
Michigan. According to Kelsall, deer weight-load-on-track (hoof only)
varies between 431 and 1,124 gm./cm.^2. However, Verme stated that
his compaction gauge (with a weight load of about 211 gm./cm.^2,
described earlier in this paper) sank in virtually the same amount in
snow as did deer. Under the snow conditions in our study area, we found
that the same type of compaction gauge generally penetrated to a depth
within a half inch of that to which deer were sinking. On this basis,
it seems reasonable to suggest that a deer in snow is supported by more
of its foot than just the hoof, and that the actual
weight-load-on-track of deer in snow is about 211 gm./cm.^2.
For wolves, this measure varies from 89 to 103 gm./cm.^2 (Foromozov
1946). This means that for the same amount of force applied during
running, a wolf would have twice as much support as a deer. It also
means that in deep snow a walking wolf generally is much less
restricted than a walking deer. Late in February 1969, for example,
when deer were seriously limited in their ability to travel, wolves
were able to travel widely (Mech _et al._, p. 1).
Even though wolves have much greater support than deer, when running
they still sink into the snow almost as much as deer under most
conditions, probably because both run with such force that snow usually
offers little support. Nevertheless, with extremely deep snow, the
difference in support factor between wolves and deer could become
critical, and this is probably what happened during February and March
1969. With deer seriously restrained by the deep snow, even a slight
advantage in favor of the wolf could increase hunting success. A high
snow density during that period would accentuate this advantage. This
is because until the snow becomes dense enough to hold a running deer,
each increase in density would further the advantage of the wolf, which
would require only half the density to support it, while it would
hinder the deer.
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