Knight found that the stimulus which influences the root to turn
downward is the force of gravity. The reaction of the root in response
to this stimulus is geotropism, a turning influenced by the earth. This
term is applied to the growth movements of plants influenced by the
earth with regard to direction. While the motor zone lies back of the
root-tip, the latter receives the stimulus and is the perceptive zone.
If the root-tip is cut off, the root is no longer geotropic, and will
not turn downward when placed in a horizontal position. Growth toward
the earth is _progeotropism_. The lateral growth of secondary roots is
_diageotropism_.
[Illustration: Fig. 112. Germinating pea placed in a horizontal
position.]
[Illustration: Fig. 113. In 24 hours gravity has caused the root to
turn downward.
Figs. 112, 113.—Progeotropism of the pea root.]
The stem, on the other hand, which was placed in a horizontal position
has become again erect. This turning of the stem in the upward
direction takes place in the dark as well as in the light, as we can
see if we start the experiment at nightfall, or place the plant in the
dark. This upward growth of the stem is also influenced by the earth,
and therefore is a case of geotropism. The special designation in the
case of upright stems is _negative geotropism_, or _apogeotropism_, or
the stems are said to be _apogeotropic_. If we place a rapidly growing
potted plant in a horizontal position by laying the pot on its side,
the ends of the shoots will soon turn upward again when placed in a
horizontal position. Young bean plants growing in a pot began within
two hours to turn the ends of the shoots upward.
[Illustration: Fig. 114. Pumpkin seedling showing apogeotropism.
Seedling at the left placed horizontally, in 24 hours the stem has
become erect.]
Horizontal leaves and shoots can be shown to be subject to the same
influence, and are therefore _diageotropic_.
=261. Influence of light.=—Not only is light a very important
factor for plants during photosynthesis, it exerts great influence on
plant growth and movement.
[Illustration: Fig. 115. Radish seedlings grown in the dark, long,
slender, not green.]
[Illustration: Fig. 116. Radish seedlings grown in the light, shorter,
stouter, and green in color. Growth retarded by light.]
=262. Growth in the absence of light.=—Plants grown in the dark
are subject to a number of changes. The stems are often longer,
more slender and weaker since they contain a larger amount of water
in proportion to building material which the plant obtains from
carbohydrates manufactured in the light. On many plants the leaves are
very small when grown in the dark.
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