=263. Influence of light on direction of growth.=—While we are
growing seedlings, the pots or boxes of some of them should be placed
so that the plants will have a one-sided illumination. This can be
done by placing them near an open window, in a room with a one-sided
illumination, or they may be placed in a box closed on all sides but
one which is facing the window or light. In 12-24 hours, or even in a
much shorter time in some cases, the stems of the seedlings will be
directed toward the source of light. This influence exerted by the rays
of light is _heliotropism_, a turning influenced by the sun or sunlight.
[Illustration: Fig. 117. Seedling of castor-oil bean, before and after
a one-sided illumination.]
[Illustration: Fig. 118. Dark chamber with opening at one side to show
heliotropism. (After Schleichert.)]
=264. Diaheliotropism.=—Horizontal leaves and shoots are
_diaheliotropic_ as well as _diageotropic_. The general direction which
leaves assume under this influence is that of placing them with the
upper surface perpendicular to the rays of light which fall upon them.
Leaves, then, exposed to the brightly lighted sky are, in general,
horizontal. This position is taken in direct response to the stimulus
of light. The leaves of plants with a one-sided illumination, as can be
seen by trial, are turned with their upper surfaces toward the source
of light, or perpendicular to the incidence of the light rays. In this
way light overcomes for the time being the direction which growth
gives to the leaves. The so-called “sleep” of plants is of course
not sleep, though the leaves “nod,” or hang downward, in many cases.
There are many plants in which we can note this drooping of the leaves
at nightfall, and in order to prove that it is not determined by the
time of day we can resort to a well-known experiment to induce this
condition during the day. The plant which has been used to illustrate
this is the sunflower. Some of these plants, which were grown in a box,
when they were about 35 _cm_ high were covered for nearly two days,
so that the light was excluded. At midday on the second day the box
was removed, and the leaves on the covered plants are well represented
by fig. 119, which was made from one of them. The leaves of the other
plants in the box which were not covered were horizontal, as shown by
fig. 120. Now on leaving these plants, which had exhibited induced
“sleep” movements, exposed to the light they gradually assumed the
horizontal position again.
[Illustration: Fig. 119. Sunflower plant. Epinastic condition of leaves
induced during the day in darkness.]
[Illustration: Fig. 120. Sunflower plant removed from darkness, leaves
extending under influence of light (diaheliotropism.)]
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