=501. The horned liverworts= take their name from the shape of
the sporogonium. This is long, slender, cylindrical, pointed, and very
slightly curved, suggesting the shape of a minute horn. Anthoceros is
one of the most common and widely distributed species. The plant grows
on damp soil or on mud.
Anthoceros.
=502. The gametophyte.=—The gametophyte is thalloid. It is thin,
flattened, green, irregularly ribbon-shaped and branched. It lies on
the soil and is more or less crisped or wavy, or curled, the edges
nearly plane, or somewhat irregular, and with minute lobes, or notches,
especially near the growing end. The general form and branching can
be seen in fig. 279. Where the plants are much crowded the thallus is
more irregular, and often possesses numerous small lateral branches
in addition to the main lobes. Upon the under side are the slender
rhizoids, which attach to the soil. With a hand lens there can be seen
also upon the under side small dark, rounded and thickened spots, where
an alga (nostoc) is located.
Sexual Organs of Anthoceros.
=502. The sexual organs of anthoceros= differ considerably from
those of the other liverworts studied. In the first place they are
immersed in the true tissue of the thallus, i.e., they do not project
above the surface.
[Illustration: Fig. 279.
Anthoceros gracilis. _A_, several gametophytes, on which sporangia
have developed; _B_, an enlarged sporogonium, showing its elongated
character and dehiscence by two valves, leaving exposed the slender
columella on the surface of which are the spores, _C_, _D_, _E_, _F_,
elaters of various forms, _G_, spores. (After Schiffner.)]
=503. Antheridia.=—The antheridium arises from an internal cell
of the thallus, a cell just below the upper surface. This cell develops
usually a group of antheridia which lie in a cavity formed around this
cell as the thallus continues to grow. They are situated along the
middle line of the thallus, and can be seen by making a section in this
direction. The antheridia are oval or rounded, have a wall of one layer
of cells which contains the sperm cells, and each antheridium has a
slender stalk. The sperms are like those of the true liverworts.
=504. Archegonia.=—The archegonia are also borne along the
middle line of the thallus. Each one arises at an early stage in the
development of the tissue of the thallus from a superficial cell,
but the archegonium does not project above the surface. The venter
therefore which contains the egg is deep down in the thallus, the wall
of the neck is formed from cells indistinguishable from the adjoining
cells of the thallus and opens at the surface.
Sporophyte of Anthoceros.
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