We have here two liquids separated by an animal membrane, water on
the one hand which diffuses readily through the membrane, while on
the other is a solution of sugar which diffuses through the animal
membrane with difficulty. The water, therefore, not containing any
solvent, according to a general law which has been found to obtain in
such cases, diffuses more readily through the membrane into the sugar
solution, which thus increases in volume, and also becomes more dilute.
The bladder membrane is what is sometimes called a diffusion membrane,
since the diffusion currents travel through it.
=43.= In this experiment then the bulk of the sugar solution is
increased, and the liquid rises in the tube by this pressure above
the level of the water in the jar outside of the thistle tube. The
diffusion of liquids through a membrane is _osmosis_.
=44. Importance of these physical processes in plants.=—Now if
we recur to our experiment with spirogyra we find that exactly the
same processes take place. The protoplasmic membrane is the diffusion
membrane, through which the diffusion takes place. The salt solution
which is first used to bathe the threads of the plant is a stronger
solution than that of the cell-sap within the cell. Water therefore
is drawn out of the cell-sap, but the substances in solution in
the cell-sap do not readily move out. As the bulk of the cell-sap
diminishes the pressure from the outside pushes the protoplasmic
membrane away from the wall. Now when we remove the salt solution and
bathe the thread with water again, the cell-sap, being a solution of
certain substances, diffuses with more difficulty than the water, and
the diffusion current is inward, while the protoplasmic membrane moves
out against the cell wall, and turgidity again results. Also in the
experiments with salt and sugar solutions on the leaves of geranium,
on the leaves and stems of the seedlings, on the tissues and cells of
the beet and carrot, and on the root hairs of the seedlings, the same
processes take place.
These experiments not only teach us that in the protoplasmic membrane,
the cell wall, and the cell-sap of plants do we have structures which
are capable of performing these physical processes, but they also show
that these processes are of the utmost importance to the plant; not
only in giving the plant the power to take up solutions of nutriment
from the soil, but they serve also other purposes, as we shall see
later.
FOOTNOTE:
Public-domain text, read in full here on John Shaqi.
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