=746. Influence of foliage leaves on the form of the stem.=—The
marked effect which foliage has upon the aspect of the plant or upon
the landscape is evident to all observers. Perhaps it is usual to
look upon the stem as having been developed for the display of the
foliage without taking into account the possibility that the foliage
may have a great influence upon the form or habit of the stem. It is
very evident, however, that the foliage exercises a great influence
on the form of the stem. For example, as trees increase in age and
size, the development of branches on the interior ceases and some of
those already formed die, since the dense foliage on the periphery
of the trees cuts off the necessary light stimulus. The tree,
therefore, possesses fewer branches and a more open interior. In the
forest also, the dense foliage above makes possible the shapely,
clean timber trunks. Note certain trees where by accident, or by
design, the terminal foliage-bearing branches have been removed that
foliage-bearing branches may arise in the interior of the tree system.
Without foliage leaves the stems of green plants would develop a very
different habit from what they do. This development could take place in
three different directions under the influence of light: (1) The light
stimulus would induce profuse branching, so that there would be many
small branches. (2) The stem would develop fewer branches, but they
would be flattened. (3) Massive trunks with but few or no branches. In
fact, all these forms are found in certain green stems which do not
bear leaves. An example of the first is found in asparagus with its
numerous crowded slender branches. But such forms in our climate are
rare, since foliage leaves are more efficient. The second and third
forms are found among cacti, which usually grow in dry regions under
conditions which would be fatal to ordinary thin foliage leaves.
=747. Relation of foliage leaves to the stem.=—In the study
of the position of the leaves on the stem we observe two important
modes of distribution: (1) the distribution along the individual
stem or branch which bears them, usually classed under the head of
_Phyllotaxy_; (2) the distribution of the leaves with reference to the
plant as a whole.
=748. Phyllotaxy, or arrangement of leaves.=—In examining buds on
the winter shoots of woody plants, we cannot fail to be impressed with
some peculiarities in the arrangement of these members on the stem of
the plant.
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