In the horse-chestnut, as we have already observed, the leaves are in
pairs, each one of the pair standing opposite its partner, while the
pair just below or above stand across the stem at right angles to the
position of the former pair. In other cases (the common bed-straw) the
leaves are in whorls, that is, several stand at the same level on the
axis, distributed around the stem. By far the larger number of plants
have their leaves arranged alternately. A simple example of alternate
leaves is presented by the elm, where the leaves stand successively on
alternate sides of the stem, so that the distance from one leaf to the
next, as one would measure around the stem, is exactly one half the
distance around the stem. This arrangement is one half, or the angle of
divergence of one leaf from the next is one half. In the case of the
sedges the angle of divergence is less, that is one-third.
By far the larger number of those plants which have the alternate
arrangement have the leaves set at an angle of divergence represented
by the fraction two fifths. Other angles of divergence have been
discovered, and much stress has been laid on what is termed a law in
the growth of the stem with reference to the position which the leaves
occupy. Singularly by adding together the numerators and denominators
of the last two fractions gives the next higher angle of divergence.
Example:
1 + 2 3 2 + 3 5
—- —- = —-; —- —- = —-;
3 + 5 8 5 + 8 13
and so on. There are, however, numerous exceptions to this regular
arrangement, which have caused some to question the importance of any
theory like that of the “spiral theory” of growth propounded by Goethe
and others of his time.
=749. Adaptation in leaf arrangement.=—As a result, however, of
one arrangement or another we see a beautiful adaptation of the plant
parts to environment, or the influence which environment, especially
light, has had on the arrangement of the leaves and branches of the
plant. Access to light and air are of the greatest importance to
green plants, and one cannot fail to be profoundly impressed with the
workings of the natural laws in obedience to which the great variety of
plants have worked out this adaptation in manifold ways.
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