=65. “Root pressure” or exudation pressure.=—It is a very common
thing to note, when certain shrubs or vines are pruned in the spring,
the exudation of a watery fluid from the cut surfaces. In the case of
the grape vine this has been known to continue for a number of days,
and in some cases the amount of liquid, called “sap,” which escapes is
considerable. In many cases it is directly traceable to the activity
of the roots, or root hairs, in the absorption of water from the soil.
For this reason the term _root pressure_ has been used to denote the
force exerted in supplying the water from the soil. But there are some
who object to the use of this term “root pressure.” The principal
objection is that the pressure which brings about the phenomenon
known as “bleeding” by plants is not present in the roots alone. This
pressure exists under certain conditions in all parts of the plant. The
term exudation pressure has been proposed in lieu of root pressure. It
should be remembered that the movement of water in the plant is started
by the pressure which exists in the root. If the term “root pressure”
is used, it should be borne clearly in mind that it does not express
the phenomenon exactly in all cases.
=Root pressure may be measured.=—It is possible to measure
not only the amount of water which the roots will raise in a given
time, but also to measure the force exerted by the roots during root
pressure. It has been found that root pressure in the case of the
nettle is sufficient to hold a column of water about 4.5 meters (15
ft.) high (Vines), while the root pressure of the vine (Hales, 1721)
will hold a column of water about 10 meters (36.5 ft.) high, and the
birch (Betula lutea) (Clark, 1873) has a root pressure sufficient to
hold a column of water about 25 meters (84.7 ft.) high.
=66. Experiment to demonstrate root pressure.=—By a very simple
method this lifting of water by root pressure is shown. During the
summer season plants in the open may be used if it is preferred, but
plants grown in pots are also very serviceable, and one may use a
potted begonia or balsam, the latter being especially useful. The
plants are usually convenient to obtain from the greenhouses, to
illustrate this phenomenon. The stem is cut off rather close to the
soil and a long glass tube is attached to the cut end of the stem,
still connected with the roots, by the use of rubber tubing, as shown
in figure 45, and a very small quantity of water may be poured in to
moisten the cut end of the stem. In a few minutes the water begins to
rise in the glass tube. In some cases it rises quite rapidly, so that
the column of water can readily be seen to extend higher and higher up
in the tube when observed at quite short intervals. (To measure the
force of root pressure is rather difficult for elementary work. To
measure it see Ganong, Plant Physiology, pp. 67, 68, or some other book
for advanced work.)
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