=849.= In some of the tropical orchids the pollinia are set free
when the insect touches a certain part of the flower, and are thrown
in such a way that the disk of the pollinium strikes the insect’s head
and stands upright. By the time the insect reaches another flower the
pollinium has bent downward sufficiently to strike against the stigma
when the insect alights on the labellum. In the mountains of North
Carolina I have seen a beautiful little orchid, in which, if one
touches a certain part of the flower with a lead-pencil or other
suitable object, the pollinium is set free suddenly, turns a complete
somersault in the air, and lands with the disk sticking to the pencil.
Many of the orchids grown in conservatories can be used to demonstrate
some of these peculiar mechanisms.
[Illustration: Fig. 464. Spray of leaves and flowers of cytisus.]
[Illustration: Fig. 465. Flower of cytisus grown in conservatory. Same
flower scattering pollen.]
=860. Pollination of the canna.=—In the study of some of the
marvellous adaptations of flowers for cross pollination one is led to
inquire if, after all, plants are not intelligent beings, instead of
mere automatons which respond to various sorts of stimuli. No plant
has puzzled me so much in this respect as the canna, and any one will
be well repaid for a study of recently opened flowers, even though it
may be necessary to rise early in the morning to unravel the mystery,
before bees or the wind have irritated the labellum. The canna flower
is a bewildering maze of petals and petal-like members. The calyx
is green, adherent to the ovary, and the limb divides into three,
lanceolate lobes. The petals are obovate and spreading, while the
stamens have all changed to petal-like members, called _staminodia_.
Only one still shows its stamen origin, since the anther is seen at one
side, while the filament is expanded laterally and upwards to form the
_staminodium_.
[Illustration: Fig. 466. Spartium, showing the dusting of the pollen
through the opening keels on the under side of an insect. (From Kerner
and Oliver.)]
=861.= The ovary has three locules, and the three styles are
usually united into a long, thin, strap-shaped style, as seen in the
figure, though in some cases three, nearly distinct, filamentous styles
are present. The end of this strap-shaped style has a peculiar curve on
one side, the outline being sometimes like a long narrow letter S. It
is on the end of this style, and along the crest of this curve, that
the stigmatic surface lies, so that the pollen must be deposited on the
stigmatic end or margin in order that fertilization may take place.
[Illustration: Fig. 467. Cypripedium.]
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