[Illustration: Fig. 468. Section of flower of cypripedium. _st_,
stigma; _a_, at the left stamen. The insect enters the labellum at
the center, passes under and against the stigma, and out through the
opening _b_, where it rubs against the pollen. In passing through
another flower this pollen is rubbed off on the stigma.]
=862.= If we open carefully canna flower buds which are nearly
ready to open naturally, by unwrapping the folded petals and
staminodia, we shall see the anther-bearing staminodium is so wrapped
around the flattened style that the anther lies closely pressed against
the face of the style, near the margin _opposite that on which the
stigma lies_.
[Illustration: Fig. 469. Epipactis with portion of perianth removed
to show details. _l_, labellum; _st_, stigma; _r_, rostellum; _p_,
pollinium. When the insect approaches the flower its head strikes the
disk of the pollinium and pulls the pollinium out. At this time the
pollinium stands up out of the way of the stigma. By the time the
insect moves to another flower the pollinia have moved downward so that
they are in position to strike the stigma and leave the pollen. At the
right is the head of a bee, with two pollinia (_a_) attached.]
[Illustration: Fig. 470. Canna flowers with the perianth removed to
show the depositing of the pollen on the style by the stamen.]
=863.= The walls of the anther locules which lie against the style
become changed to a sticky substance for their entire length, so that
they cling firmly to the surface of the style and also to the mass of
pollen within the locules. The result is that when the flower opens,
and this staminodium unwraps itself from the embrace of the style,
the mass of pollen is left there deposited, while the empty anther is
turned around to one side.
=668.= Why does the flower deposit its own pollen on the style?
Some have regarded this as the act of pollination, and have concluded,
therefore, that cannas are necessarily self pollinated, and that cross
pollination does not take place. But why is there such evident care to
deposit the pollen on the side of the style away from the stigmatic
margin? If we visit the cannas some morning, when a number of the
flowers have just opened, and the bumblebees are humming around seeking
for nectar, we may be able to unlock the secret.
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