=88. Root pressure may exceed transpiration.=—If we cover small
actively growing plants, such as the pea, corn, wheat, bean, etc.,
with a bell jar, and place them in the sunlight where the temperature
is suitable for growth, in a few hours, if conditions are favorable,
we shall see that there are drops of water standing out on the margins
of the leaves. These drops of water have exuded through the ordinary
stomata, or in other cases what are called water stomata, through the
influence of root pressure. The plant being covered by the glass jar,
the air soon becomes saturated with moisture and transpiration is
checked. Root pressure still goes on, however, and the result is shown
in the exuding drops. Root pressure is here in excess of transpiration.
This phenomenon is often to be observed during the summer season in the
case of low-growing plants. During the bright warm day transpiration
equals, or may be in excess of, root pressure, and the leaves are
consequently flaccid. As nightfall comes on the air becomes more
moist, and the conditions of light are such also that transpiration
is lessened. Root pressure, however, is still active because the soil
is still warm. In these cases drops of water may be seen exuding from
the margins of the leaves due to the excess of root pressure over
transpiration. Were it not for this provision for the escape of the
excess of water raised by root pressure, serious injury by lesions, as
a result of the great pressure, might result. The plant is thus to some
extent a self-regulatory piece of apparatus so far as root pressure and
transpiration are concerned.
=89. Injuries caused by excessive root pressure.=—Some varieties
of tomatoes when grown in poorly lighted and poorly ventilated
greenhouses suffer serious injury through lesions of the tissues.
This is brought about by the cells at certain parts becoming charged
so full with water through the activity of root pressure and lessened
transpiration, assisted also probably by an accumulation of certain
acids in the cell-sap which cannot be got rid of by transpiration.
Under these conditions some of the cells here swell out, forming
extensive cushions, and the cell walls become so weakened that they
burst. It is possible to imitate the excess of root pressure in the
case of some plants by connecting the stems with a system of water
pressure, when very quickly the drops of water will begin to exude from
the margins of the leaves.
[Illustration: Fig. 56.
The roots are lifting more water into the plant than can be given off
in the form of water vapor, so it is pressed out in drops. From “First
Studies Plant Life.”]
=90.= It should be stated that in reality there is no difference
between transpiration and evaporation, if we bear in mind that
evaporation takes place more slowly from living plants than from dead
ones, or from an equal surface of water.
Public-domain text, read in full here on John Shaqi.
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