=97.= We thus see that instead of the liquids passing through the
entire stem they are confined to definite courses. Now that we have
discovered the path of the upward movement of water in the stem, we are
curious to see what the structure of these definite portions of the
stem is.
[Illustration: Fig. 58.
Xylem portion of bundle. Cambium portion of bundle. Bast portion of
bundle.
Section of vascular bundle of sunflower stem.]
=98. Structure of the fibrovascular bundles.=—We should now make
quite thin cross-sections, either free hand and mount in water for
microscopic examination, or they may be made with a microtome and
mounted in Canada balsam, and in this condition will answer for future
study. To illustrate the structure of the bundle in one type we may
take the stem of the castor-oil bean. On examining these cross-sections
we see that there are groups of cells which are denser than the ground
tissue. These groups correspond to the colored areas in the former
experiments, and are the vascular bundles cut across. These groups are
somewhat oval in outline, with the pointed end directed toward the
center of the stem. If we look at the section as a whole we see that
there is a narrow continuous ring[7] of small cells situated at the
same distance from the center of the stem as the middle part of the
bundles, and that it divides the bundles into two groups of cells.
=99. Woody portion of the bundle.=—In that portion of the bundle
on the inside of the ring, i.e., toward the “pith,” we note large,
circular, or angular cavities. The walls of these cells are quite thick
and woody. They are therefore called wood cells, and because they
are continuous with cells above and below them in the stem in such a
way that long tubes are formed, they are called woody vessels. Mixed
in with these are smaller cells, some of which also have thick walls
and are wood cells. Some of these cells may have thin walls. This
is the case with all when they are young, and they are then classed
with the fundamental tissue or soft tissue (parenchyma). This part of
the bundle, since it contains woody vessels and fibres, is the _wood
portion_ of the bundle, or technically the _xylem_.
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