=103. Vessels or ducts.=—One way in which the cells in side view
differ greatly from an end view, in a cross-section in the bundle, is
that they are much longer in the direction of the axis of the stem. The
cells have become elongated greatly. If we search for the place where
two of these large cells with spiral, or ladder-like, markings meet end
to end, we see that the wall which formerly separated the cells has
nearly or quite disappeared. In other words the two cells have now an
open communication at the ends. This is so for long distances in the
stem, so that long columns of these large cells form tubes or vessels
through which the water rises in the stems of plants.
=104.= In the bast portion of the bundle we detect the cells of
the bast fibers by their thick walls. They are very much elongated and
the ends taper out to thin points so that they overlap. In this way
they serve to strengthen the stem.
=105. Sieve tubes.=—Lying near the bast cells, usually toward
the cambium, are elongated cells standing end to end, with delicate
markings on their cross walls which appear like finely punctured plates
or sieves. The protoplasm in such cells is usually quite distinct, and
sometimes contracted away from the side walls, but attached to the
cross walls, and this aids in the detection of the sieve tubes (fig.
59.) The granular appearance which these plates present is caused by
minute perforations through the wall so that there is a communication
between the cells. The tubes thus formed are therefore called sieve
tubes and they extend for long distances through the tube so that there
is communication throughout the entire length of the stem. (The
function of the sieve tubes is supposed to be that for the downward
transportation of substances elaborated in the leaves.)
=106.= If we section in like manner the stem of the sunflower we
shall see similar bundles, but the number is greater than eight. In
the garden balsam the number is from four to six in an ordinary stem
3-4_mm_ diameter. Here we can see quite well the origin of the vascular
bundle. Between the larger bundles we can see especially in free-hand
sections of stems through which a colored solution has been lifted by
transpiration, as in our former experiments, small groups of the minute
cells in the cambial ring which are colored. These groups of cells
which form strands running through the stem are _pro-cambium strands_.
The cells divide and increase just like the cambium cells, and the
older ones thrown off on either side change, those toward the center
of the stem to wood vessels and fibers, and those on the outer side to
bast cells and sieve tubes.
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