Inulin, a carbohydrate closely related to starch, is stored up for
food in solution in many composite plants, as in the artichoke, the
root tuber of dahlia, etc. When used for food by the growing plant
it is converted into glucose by an enzyme, inulase. Make a section
of a portion of a dahlia tuber or artichoke and treat with alcohol.
The inulin is precipitated into sphæro crystals. (See also paragraphs
156-161 and 216_b_.)
=218.= Then there are certain fungi which feed on starch or other
organic substances whether in the host or not, which excrete certain
enzymes to dissolve the starch, etc., to bring it into a soluble
form before they can absorb it as food. Such a process is a sort of
_extracellular digestion_, i.e., the organism excretes the enzyme and
digests the solid outside, since it cannot take the food within its
cells in the solid form. To a certain degree the higher plants perform
also extracellular digestion in the action of root hair excretion on
insoluble substances, and in the case of the humus saprophytes. But for
them soluble food is largely prepared by the action of acids, etc., in
the soil or water, or by the work of fungi and bacteria as described in
Chapter 9.
=219. Assimilation.=—In plant physiology the term assimilation
has been chiefly used for the process of carbon dioxide assimilation
(= photosynthesis). Some objections have been raised against the use
of assimilation here as one of the life processes of the plant, since
its inception stages are due to the combined action of light, an
external factor, and chlorophyll in the plant along with the living
chloroplastid. So long, however, as it is not known that this process
can take place without the aid of the living plant, it does not seem
proper to deny that it is altogether not a process of assimilation. It
is not necessary to restrict the term assimilation to the formation
of new living matter in the plant cell; it can be applied also to
the synthetic processes in the formation of carbohydrates, proteids,
etc., and called synthetic assimilation. The sun supplies the energy,
which is absorbed by the chlorophyll, for splitting up the carbonic
acid, and the living chloroplast then assimilates by a synthetic
process the carbon, hydrogen, and oxygen. This process then can be
called _photosynthetic assimilation_. The nitrite and nitrate bacteria
derive energy in the process of nitrification, which enables them
to assimilate CO₂ from the air, and this is called _chemosynthetic
assimilation_. The inorganic material in the form of mineral salts,
nitrates, etc., absorbed by the root, and carried up to the leaves,
here meets with the carbohydrates manufactured in the leaf. Under
the influence of the protoplasm synthesis takes place, and proteids
and other organic compounds are built up by the union of the salts,
nitrates, etc., with the carbohydrates. This is also a process of
synthetic assimilation. These are afterward stored as food, or
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