Elements of Agricultural ChemistryAnderson, Thomas
Science
Elements of Agricultural Chemistry
Anderson, Thomas
Agricultural chemistry
_Changes occurring during Germination._--When a seed is placed in the
soil under favourable circumstances, it becomes the seat of an important
and remarkable series of chemical changes, which result in the
production of the young plant. Experiment and observation have shown
that heat, moisture, and air, are necessary to the production of these
changes, and though probably not absolutely essential, the absence of
light is favourable in the early stages. The temperature required for
germination varies greatly in different seeds, some germinating readily
at a few degrees above the freezing point, and others requiring a
tolerably high temperature. The rapidity with which it takes place
appears to increase with the temperature; but this is true only within
very narrow limits, for beyond a certain point heat is injurious, and
when it exceeds 120 deg. or 130 deg. Fahrenheit, entirely prevents the process.
The presence of oxygen is also essential, for it has been shown that if
seeds are placed in a soil exposed to an atmosphere deprived of that
element, or if they be buried so deep that the air does not reach them,
they may lie without change for an unlimited period; but so soon as they
are exposed to the air, germination immediately commences. Illustrations
of this fact are frequently observed where earth from a considerable
depth has been thrown up to the surface, when it often becomes covered
with plants not usually seen in the neighbourhood, which have sprung
from buried seeds. When all the necessary conditions for germination are
fulfilled, the seed absorbs moisture, swells up, and sends out a shoot
which rises to the surface, and a radicle which descends--the one
destined to develop the leaves, the other the roots, by which the plant
is afterwards to derive its nutriment from the air and the soil. But
until these organs are properly developed, the plant is dependent on the
matters contained in the seed itself. These substances are mostly
insoluble, but are brought into solution by the atmospheric oxygen
acting upon the gluten, and converting it into a soluble substance
called diastase, which in its turn reacts upon the starch, converting it
first into dextrine, and then into cellulose, and the latter is finally
deposited in the form of organised cells, and produces the first little
shoot of the plant. At the first moment of germination, the oxygen
absorbed appears simply to oxidize the constituents of the seed, but
this condition exists only for a very limited period, and is soon
followed by the evolution of carbonic acid, water being at the same time
formed from the organic constituents of the seed, which gradually
diminishes in weight. The amount of this diminution is different with
different plants, but always considerable. Boussingault found that the
loss of dry substance in the pea amounted in 26 days to 52 per cent, and
in wheat to 57 per cent in 51 days. Against this, of course, is to be
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