Elements of Structural and Systematic Botany: For High Schools and Elementary College CoursesCampbell, Douglas Houghton
Science
Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses
Campbell, Douglas Houghton
Botany -- Textbooks
Running around the outside of the ripe spore case is a single row of
cells (_an._), differing from the others in shape, and having their
inner walls thickened. Near the bottom, two (sometimes four) of
these cells are wider than the others, and their walls are more
strongly thickened. It is at this place (_st._) that the spore case
opens. When the ripe sporangium becomes dry, the ring of thickened
cells (_an._) contracts more strongly than the others, and acts like
a spring pulling the sporangium open and shaking out the spores,
which germinate readily under favorable conditions, and form after a
time the sexual plants (prothallia).
The roots of the sporophyte arise in large numbers, the youngest being
always nearest the growing point of the stem or larger roots (Fig. 67,
_C_). The growing roots are pointed at the end which is also
light-colored, the older parts becoming dark brown. A cross-section of
the older portions shows a dark-brown ground tissue with a central,
light-colored, circular, fibro-vascular bundle (Fig. 68, _F_). Growing
from its outer surface are numerous brown root hairs (_r_).
When magnified the walls of all the outer cells (epidermis and
ground tissue) are found to be dark-colored but not very thick, and
the cells are usually filled with starch. There is a bundle sheath
of much-flattened cells separating the fibro-vascular bundle from
the ground tissue. The bundle (Fig. 68, _G_) shows a band of
tracheary tissue in the centre surrounded by colorless cells, all
about alike.
All of the organs of the fern grow from a definite apical cell, but
it is difficult to study except in the root.
Selecting a fresh, pretty large root, a series of thin longitudinal
sections should be made either holding the root directly in the
fingers or placing it between pieces of pith. In order to avoid
drying of the sections, as is indeed true in cutting any delicate
tissue, it is a good plan to wet the blade of the razor. If the
section has passed through the apex, it will show the structure
shown in Figure 68, _D_. The apical cell (_a_) is large and
distinct, irregularly triangular in outline. It is really a
triangular pyramid (tetrahedron) with the base upward, which is
shown by making a series of cross-sections through the root tip, and
comparing them with the longitudinal sections. The cross-section of
the apical cell (Fig. _L_) appears also triangular, showing all its
faces to be triangles. Regular series of segments are cut off in
succession from each of the four faces of the apical cell. These
segments undergo regular divisions also, so that very early a
differentiation of the tissues is evident, and the three tissue
systems (epidermal, ground, and fibro-vascular) may be traced
almost to the apex of the root (68, _D_). From the outer series of
segments is derived the peculiar structure (root cap) covering the
delicate growing point and protecting it from injury.
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