Elements of Structural and Systematic Botany: For High Schools and Elementary College CoursesCampbell, Douglas Houghton
Science
Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses
Campbell, Douglas Houghton
Botany -- Textbooks
After the archegonia are fertilized the outer parts of the ovule
become hard and brown, and serve to protect the embryo plant, which
reaches a considerable size before the sporangium falls off. As the
walls of the ovule harden, the carpel or leaf bearing it undergoes a
similar change, becoming extremely hard and woody, and as each one
ends in a sharp spine, and they are tightly packed together, it is
almost impossible to separate them. The ripe cone (Fig. 75, _A_)
remains closed during the winter, but in the spring, about the time
the flowers are mature, the scales open spontaneously and discharge
the ripened ovules, now called seeds. Each seed (_E_, _s_) is
surrounded by a membranous envelope derived from the scale to which it
is attached, which becomes easily separated from the seed. The opening
of the cones is caused by drying, and if a number of ripe cones are
gathered in the winter or early spring, and allowed to dry in an
ordinary room, they will in a day or two open, often with a sharp,
crackling sound, and scatter the ripe seeds.
A section of a ripe seed (_F_) shows the embryo (_em._) surrounded by
a dense, white, starch-bearing tissue derived from the prothallium
cells, and called the "endosperm." This fills up the whole seed which
is surrounded by the hardened shell derived from the integument and
wall of the ovule. The embryo is elongated with a circle of small
leaves at the end away from the opening of the ovule toward which is
directed the root of the embryo.
The seed may remain unchanged for months, or even years, without
losing its vitality, but if the proper conditions are provided, the
embryo will develop into a new plant. To follow the further growth of
the embryo, the ripe seeds should be planted in good soil and kept
moderately warm and moist. At the end of a week or two some of the
seeds will probably have sprouted. The seed absorbs water, and the
protoplasm of the embryo renews its activity, beginning to feed upon
the nourishing substances in the cells of the endosperm. The embryo
rapidly increases in length, and the root pushes out of the seed
growing rapidly downward and fastening itself in the soil (_G_, _r_).
Cutting the seed lengthwise we find that the leaves have increased
much in length and become green (one of the few cases where
chlorophyll is formed in the absence of light). As these leaves
(called "cotyledons" or seed leaves) increase in length, they
gradually withdraw from the seed whose contents they have exhausted,
and the young plant enters upon an independent existence.
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