Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10Various
Science
Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10
Various
Encyclopedias and dictionaries
This class shows much greater diversity of organization than any
other, and the classifications proposed by recent writers, such as E.
Haeckel, O. Jaekel and F.A. Bather, start from such different points
of view that no discussion of them can be attempted here. Following
the narrative given above, we recognize a primitive
group--Amphoridea--represented by _Aristocystis_ (fig. 8). From this
are derived the orders Diploporita (fig. 9) and Rhombifera (fig. 10)
and the class Edrioasteroidea, all which have already been described
as steps in the evolution of the phylum. But there were also
side-branches leading nowhere, and therefore placed in separate
orders--Aporita and Carpoidea.
Order 1. _Amphoridea._--Radial symmetry has affected neither
food-grooves nor thecal plates; nor, probably, nerves, ambulacral
vessels, nor gonads. Canals or folds when present in the stereom are
irregular. Families: Aristocystidae (fig. 8); Eocystidae.
Order 2. _Carpoidea._--Theca compressed in the oro-anal plane and a
bilateral symmetry thus induced, affecting the food-grooves and,
usually, the thecal plates and stem. Food-grooves in part epithecal
and may be continued on one or two exothecal processes. No pores or
folds in the stereom. Families: Anomalocystidae, Dendrocystidae. These
correspond to Jaekel's Carpoidea Heterostelea; he also includes, as
Eustelea, our Comarocystidae and Malocystidae.
Order 3. _Rhombifera._--Radial symmetry affects the food-grooves and,
in the more advanced families, the thecal plates; probably also the
nerves and ambulacral vessels, but not the gonads. The food-grooves
are exothecal, i.e. are stretched out from the theca on jointed
skeletal processes (brachioles). These either are close to the mouth
or are removed from it upon a series of ambulacral or sub-ambulacral
plates not derived immediately from thecal plates, or are separated
from the oral centre by hypothecal passages passing beneath terminal
plates. The stereom and stroma become arranged in folds and strands at
right angles to the sutures of the thecal plates; in higher forms the
stereom-folds are in part specialized as pectini-rhombs. Families:
Echinosphaeridae; Comarocystidae; Macrocystellidae; Tiaracrinidae;
Malocystidae; Glyptocystidae, with sub-famm. Echinoencrininae,
Callocystinae, Glyptocystinae, of which examples are _Cheirocrinus_
(fig. 10) and _Cystoblastus_ from which Jaekel deduces the blastoids;
Caryocrinidae.
Order 4. _Aporita._--Pentamerous symmetry affects the food-grooves and
thecal plates; probably also the nerves and ambulacral vessels, but
not the gonads. Food-grooves exothecal and circumoral. The stereom
shows no trace of canals, folds, rhombs or diplopores. Family:
Cryptocrinidae.
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