Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10Various
Science
Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10
Various
Encyclopedias and dictionaries
No existing classification of the Asterida is satisfactory even for
the recent forms, still less when the older fossils are considered. A
separation of the latter as Palasterida, because of their alternating
ambulacrals, from the recent Euasterida with opposite ambulacrals, is
now discarded and an attempt made to arrange the Palasterida in
divisions originally established for Euasterida. Those divisions fall
under three schemes. C. Viguier has divided the starfish into:
_Astéries ambulacraires_, with plates of ambulacral origin prominent
in the mouth-skeleton, pedicellariae stalked, and straight or crossed,
podial pores usually quadriserial; _Astéries adambulacraires_, with
adambulacrals prominent in the mouth-skeleton, pedicellariae sessile,
and forcipiform or valvular, podial pores usually biserial. Perrier,
at first laying greater stress on the nature of the pedicellariae and
afterwards on the form of the mouth-skeleton, has gradually perfected
a scheme of five orders: (1) _Forcipulata_, with pedicellariae
stalked, and straight or crossed; (2) _Spinulosa_, with pedicellariae
sessile and forcipiform; (3) _Velata_, with membraniferous spines; (4)
_Paxillosa_, pedicellariae represented by an ossicle of the test and
the spines covering it, the whole forming a paxilla; (5) _Valvata_ or
_Granulosa_, with pedicellariae sessile and valvular or salt-cellar
shaped. A more widely accepted scheme is that of W.P. Sladen, who
divided the Euasterida into two orders; (1) _Phanerozonia_, with
marginals large and highly developed, the supero-marginals and
infero-marginals contiguous, with papulae confined to the dorsal
surface, with ambulacrals well spaced and usually broad, adambulacrals
prominent in the mouth-skeleton, with pedicellariae sessile; (2)
_Cryptozonia_, with marginals inconspicuous and somewhat atrophied in
the adult, the supero-marginals separated from the infero-marginals by
intercalated plates, with papulae distributed over the whole body,
with ambulacrals crowded and narrow, either ambulacrals or
adambulacrals prominent in the mouth-skeleton, with pedicellariae
stalked or sessile.
We give here a list of the families separated into Sladen's orders and
grouped under Perrier's divisions, extinct families being marked [+].
[Illustration: FIG. 16.--Section across the arm-skeleton of a
Phanerozonate Asterid, _Astropecten_.
a, Ambulacral plates.
b, Adambulacral plates.
c and d, Inferior and superior lateral plates.
e, Dorsal plates with paxillae. Certain supra-ambulacral plates,
which also exist, are not shown.]
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