Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10Various
Science
Encyclopaedia Britannica, 11th Edition, "Echinoderma" to "Edward, prince of Wales": Volume 8, Slice 10
Various
Encyclopedias and dictionaries
the rocks or under stones, or at times in empty Mollusc or Echinid
shells. They are occasionally used by fishermen for bait.
[Illustration: FIG. 2.--Female _Bonellia viridis_, Rol. Opened along the
left side.
a, Proboscis cut short.
b, Bristle passing through the mouth into the pharynx.
c, Coiled intestine.
d, Anal tufts or vesicles.
e, Ventral nerve cord.
f, Ovary borne on ventral vessel running parallel with e.
g, Position of anus.
h, Position of external opening of nephridium.
i, Nephridium--the line points towards, but does not reach, the
internal opening.]
_Anatomy_ (fig. 2).--A thin cuticle covers the epidermis, which
contains mucus-secreting glands. Beneath the epidermis is a layer of
circular muscles, then a layer of longitudinal, and finally in some
cases a layer of oblique muscle-fibres. The inner face of this
muscular skin is lined by a layer of epithelium. The coelomic
body-cavity is spacious. It does not extend into the proboscis, which
is a solid organ traversed by the nervous and vascular rings, but
otherwise largely built up of muscle fibres and connective tissue.
Many sense-cells lie in the epidermis. The ciliated ventral groove of
the proboscis leads at its base into the simple mouth, which gives
access to the thin-walled alimentary canal. This is longer than the
body, and to tuck it away it is looped from side to side. The loops
are supported by strands of connective tissue, which in some species
are united so as to form a dorsal mesentery, whilst traces of a
ventral mesentery are met with anteriorly and posteriorly (H.L.
Jameson, _Zool. Jahrb. Anat._, 1899). The alimentary canal is
divisible into fore-gut, mid-gut and hind-gut, and the first-named can
be further divided into pharynx, oesophagus, gizzard and crop, mainly
on histological grounds. The mid-gut is characterized by the presence
of a ciliated groove, from which arises the collateral intestine or
siphon, a second tube which rejoins the alimentary canal lower down.
Similar collateral intestines are familiar in the Echinids and certain
Polychaets (Capitellidae). The rectum receives the openings of a pair
of very characteristic organs, the anal vesicles. Each consists of a
branching tube, the tips of whose twigs terminate in minute ciliated
funnels. The anal vesicles are thought to be excretory; whether this
be so or not, they undoubtedly have some influence on the amount of
fluid found in the coelom. The coelomic fluid contains as a rule both
amoeboid and rounded corpuscles, and, when ripe, the products of the
gonads. A closed system of vessels, usually called the vascular
system, is present. There are, however, no capillaries connected with
this, and it is confined to certain portions of the body. It can
possess few of the functions usually associated with a vascular
system, and its main use is probably to assist in the expansion of the
proboscis.
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