In the Hippoidea there is generally the full series of 44 teeth, but the first premolar is often deciduous or wanting in the lower or in both jaws. The incisors are chisel-shaped, and the canines tend to become isolated so as in the now specialized forms to occupy nearly the middle of a longer or shorter gap between the incisors and premolars. In the upper molars the two outer columns of the primitive tubercular molar coalesce to form an outer wall, from which proceed two crescentic transverse crests; the connexion between the crests and the wall being imperfect or slight, and the crests themselves sometimes tubercular. Each of the lower molars carries two crescentic ridges. The number of toes ranges from four to one in the fore-foot, and from three to one in the hind-foot. The paroccipital, postglenoid and post-tympanic processes of the skull are large, and the latter always distinct. Normally there are no traces of horn-cores. The calcaneum lacks the facet for the fibula found in the Titanotheroidea. In the earlier _Equidae_ the teeth were short-crowned, with the premolars simpler than the molars; but there is a gradual tendency to an increase in the height of the crowns of the teeth, accompanied by increasing complexity of structure and the filling up of the hollows with cement. Similarly the gap on each side of the canine tooth in each jaw continues to increase in length; while in all the later forms the orbit is surrounded by a ring of bone. A third modification is the increasing length of limb (as well as in general bodily size), accompanied by a gradual reduction in the number of toes from three or four to one. [Illustration: FIG. 1.--a, Side view of second upper molar tooth of _Anchitherium_ (brachyodont form); b, corresponding tooth of horse (hypsidont form).] All the existing members of the family, such as the domesticated horse (_Equus caballus_) and its wild or half-wild relatives, the asses and the zebras, are included in the typical genus. In all these the crowns of the cheek-teeth are very tall (fig. 1, b) and only develop roots late in life; while their grinding-surfaces (fig. 2, b and c) are very complicated and have all the hollows filled with cement. The summits of the incisors are infolded, producing, when partially worn, the "mark." In the skull the orbit is surrounded by bone, and there is no distinct depression in front of the same. Each limb terminates in one large toe; the lateral digits being represented by the splint-bones, corresponding to the lateral metacarpals and metatarsals of _Hipparion_. Not unfrequently, however, the lower ends of the splint-bones carry a small expansion, representing the phalanges.