EQUILIBRIUM (from the Lat. _aequus_, equal, and _libra_, a balance), a condition of equal balance between opposite or counteracting forces. By the "sense of equilibrium" is meant the sense, or sensations, by which we have a feeling of security in standing, walking, and indeed in all the movements by which the body is carried through space. Such a feeling of security is necessary both for maintaining any posture, such as standing, or for performing any movement. If this feeling is absent or uncertain, or if there are contradictory sensations, then definite muscular movements are inefficiently or irregularly performed, and the body may stagger or fall. When we stand erect on a firm surface, like a floor, there is a feeling of resistance, due to nervous impulses reaching the brain from the soles of the feet and from the muscles of the limbs and trunk. In walking or running, these feelings of resistance seem to precede and guide the muscular movements necessary for the next step. If these are absent or perverted or deficient, as is the case in the disease known as locomotor ataxia, then, although there is no loss of the power of voluntary movement, the patient staggers in walking, especially if he is not allowed to look at his feet, or if he is blind-folded. He misses the guiding sensations that come from the limbs; and with a feeling that he is walking on a soft substance, offering little or no resistance, he staggers, and his muscular movements become irregular. Such a condition maybe artificially brought about by washing the soles of the feet with chloroform or ether. And it has been observed to exist partially after extensive destruction of the skin of the soles of the feet by burns or scalds. This shows that tactile impulses from the skin take a share in generating the guiding sensation. In the disease above mentioned, however, tactile impressions may be nearly normal, but the guiding sensation is weak and inefficient, owing to the absence of impulses from the muscles. The disease is known to depend on morbid changes in the posterior columns of the spinal cord, by which impulses are not freely transmitted upwards to the brain. These facts point to the existence of impulses coming from the muscles and tendons. It is now known that there exist peculiar spindles, in muscle, and rosettes or coils or loops of nerve fibres in close proximity to tendons. These are the end organs of the sense. The transmission of impulses gives rise to the _muscular sense_, and the guiding sensation which precedes co-ordinated muscular movements depends on these impulses. Thus from the limbs streams of nervous impulses pass to the sensorium from the skin and from muscles and tendons; these may or may not arouse consciousness, but they guide or evoke muscular movements of a co-ordinated character, more especially of the limbs.