the chromidial mass with the micronucleus, whose chief function is
reproductive.
[Illustration: FIG. 14.--Vertical section of tubulated chamber-walls,
a, a, of _Nummulites_. b, b, Marginal cord; c, cavity of chamber; d,
d, non-tubulated columns.]
Since megalospheric forms are by far the most abundant, it seems
probable that under most conditions they also give rise to
megalospheric young like themselves; and that the production of
zoospores, pairing to pass into the microspheric form, is only
occasional, and possibly seasonal. This life-history we owe to the
researches of Schaudinn and J.J. Lister.
In several species (notably _Patellina_) plastogamy, the union of the
cytoplasmic bodies without nuclear fusion, has been noted, as a
prelude to the resolution of the conjoined protoplasm into uninucleate
amoebulae.
_Calcituba_, a porcellanous type, which after forming the embryonic
chamber with its deflected pylome grows into branching stems, may fall
apart into sections, or the protoplasm may escape and break up into
small amoebulae. Of the reproduction of the simplest forms we know
little. In _Mikrogromia_ the cell undergoes fission within the test,
and on its completion the daughter-cells may emerge as biflagellate
zoospores.
The sandy shells are a very interesting series. In _Astrorhiza_ the
sand grains are loosely agglutinated, without mineral cement; they
leave numerous pores for the exit of the protoplasm, and there are no
true pylomes. In other forms the union of the grains by a calcareous
or ferruginous cement necessitates the existence of distinct pylomes.
Many of the species reproduce the varieties of form found in
calcareous tests; some are finely perforated, others not. Many of the
larger ones have their walls thickened internally and traversed by
complex passages; this structure is called _labyrinthic_ (fig. 19, g,
h). The shell of _Endothyra_, a form only known to us by its abundance
in Carboniferous and Triassic strata, is largely composed of calcite
and is sometimes perforated.
[Illustration: FIG. 15.--_Cycloclypeus_.]
It is noteworthy that though of similar habitat each species selects
its own size or sort of sand, some utilizing the siliceous spicules of
sponges. Despite the roughness of the materials, they are often so
laid as to yield a perfectly smooth inner wall; and sometimes the
outer wall may be as simple. As we can find no record of a deflected
stylopyle to the primitive chamber of the polythalamous Arenacea, it
is safe to conclude that they have no close alliance with the
Porcellanea.
_Classification._
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