The pelagic floating genera are also specially modified. Their shell is
either thin or extended many times by long slender tapering spines, and
the protoplasm outside has the same character as that of the Radiolaria
(q.v.), being differentiated into jelly containing enormous vacuoles and
traversed by reticulate strands of granular protoplasm. These coalesce
into a peripheral zone from which protrude the pseudopods, here rather
radiate than reticulate. Most genera and most species are cosmopolitan;
but local differences are often marked. Foraminifera abound in the shore
sands and the crevices of coral reefs. The membranous shelled forms
decay without leaving traces. The sandy or calcareous shells of dead
Foraminifera constitute a large proportion of littoral sand, both below
and above tide marks; and, as shown in the boring on Funafuti, enter
largely into the constituents of coral rock. They may accumulate in the
mud of the bottom to constitute Foraminiferal ooze. The source of these
shells in the latter case is double: (1) shells of bottom-dwellers
accumulate on the spot; (2) shells of dead plankton forms sink down in a
continuous shower, to form a layer at the bottom of the ocean, during
which process the spines are dissolved by the sea-water. Thus is formed
an ooze known as "Globigerina-ooze," being formed largely of that genus
and its ally _Hastigerina_; below 3000 fathoms even the tests themselves
are dissolved. Casts of their bodies in glauconite (a green ferrous
silicate, whose composition has not yet been accurately determined) are,
however, frequently left. Glauconitic casts of perforate shells, notably
_Globigerina_, have been found in Lower Cambrian (e.g. Hollybush
Sandstone), and the shells themselves in Siberian limestones of that
age. It is only when we pass into the Silurian Wenlock limestone that
sandy shells make their appearance. Above this horizon Foraminifera are
more abundant as constituents, partial or principal of calcareous rocks,
the genus _Endothyra_ being indeed almost confined to Carboniferous
beds. The genus _Fusulina_ (fig. 20) and _Saccammina_ (fig. 19) give
their names (from their respective abundance) to two limestones of the
Carboniferous series. Porcellanous shells become abundant only from the
Lias upwards. The glauconitic grains of the Greensand formations are
chiefly foraminiferal casts. Chalk is well known to consist largely of
foraminiferal shells, mostly vitreous, like the north Atlantic
globigerina ooze. In the Maestricht chalk more littoral conditions
prevailed, and we find such large-sized species as _Orbitoides_
(vitreous) and _Orbitolites_ (porcellanous; figs. 5, 6), &c. In the
Eocene Tertiaries the Calcaire Grossier of the Paris basin is mainly
composed of Miliolid forms. Nummulites occur in English beds and in the
Paris basin; but the great beds of these, forming reef-like masses of
limestone, occur farther south, extending from the Pyrenees through the
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